Culladia Moore, 1886

Culladia Moore, 1886 View in CoL

Culladia Moore, 1886: 382 .

Type species: Araxes admigratella Walker, 1863

= Araxes * Walker, 1863 b: 192. Type species: Araxes admigratella Walker, 1863

= Crambidion Mabille, 1900: 748 . Type species: Crambidion achroellum Mabille, 1900 . Viette 1990: 85 (syn.)

= Nirmaladia Rose, 1983: 172 , 175, figs. 1–7. Type species: Culladia dentilinaelis Hampson, 1919 . M. Shaffer, Nielsen & Horak 1996: 183 (syn.)

DIAGNOSIS

The genus Culladia was revised by Bleszynski (1970a). Species of Culladia exhibit small, greyish wings. Vein R5 is very short or missing as observed in a few small Crambini genera ( Bleszynski, 1970a; Léger et al., 2019). In male genitalia, the bifid sclerotized apex of the phallus is a potential apomorphy for the genus. Male genitalia exhibit the following characters: uncus and gnathos slender, with apex pointing downwards; valva slender, with costal arm at base, ventral process absent; pseudosaccus present; juxta short, with deep notch; vesica with one or more cornuti, with characteristic bifid sclerotized apex ( Bleszynski, 1970a; pers. obs.). Female genitalia exhibit the following characters: papillae anales not coalesced dorsally and ventrally; posterior apophyses 1-2 X length of tergite VIII; anterior apophyses very short; antrum often with pair of sclerites; membranous pouch branching near antrum found in several species; corpus bursae usually with long, scobinate signum ( Bleszynski, 1970a; pers. obs.)

DISTRIBUTION

Occuring in the Afrotropical, Oriental, and Australasian regions. Five species are reported from the Philippines: Culladia evae Bleszynski, 1970 , C. hastiferalis (Walker, 1865) , C. pseudoscoparia sp. n., C. suffusella Bleszynski , C. tonkinella Bleszynski, 1970 .

PHYLOGENETIC RELATIONSHIPS

Culladia is sister to the Holarctic genus Pediasia Hübner, 1825 ( Bleszynski, 1970a; Léger et al., 2019).