Aconias longisetosus, Watanabe, 2025

Aconias longisetosus sp. nov.

[New SJN: Kenaga-togari-himebachi]

( Figs. 4A–H, 77F, 78C)

Type series. Holotype: JAPAN, KPM-NK 81196 , F, Kagoshima Pref., Tokunoshima Is., Kedoku, 22. V. 2008, A. Sakai leg. Paratype: JAPAN: KPM-NK 81197 , F, Kagoshima Pref., Amamioshima Is., Yamato Vil., Oodana , 1. VI. 2007, M. Gunji leg. ; KPM-NK 81198 , 103174 , 103175 , 3 F, same locality of holotype except for 21. V. 2008, K. Watanabe leg.

Description. Female (n = 5). Body polished; covered with setae; body length 5.25–7.4 (HT: 7.1) mm.

Head 0.6–0.65 (HT: 0.63) × as long as wide in dorsal view. Clypeus 3.4–3.5 (HT: 3.45) × as wide as long; slightly convex in lateral view; sparsely punctate dorsally; smooth ventrally; punctures partly united into groove-like foveola; lower margin weakly rounded; blunt in lateral view. Face 0.38–0.4 (HT: 0.38) × as long as minimum width; slightly convex medially; matt; densely punctate medially; punctures partly united into groove-like foveola medially. Frons weakly concave above antennal sockets; irregularly or obliquely rugose except for narrowly matt along orbits. POL 1.1–1.5 (HT: 1.2) × as OD. OOL 0.95–1.4 (HT: 1.15) × as OD. Gena and occiput densely punctate. Dorsal profile of gena rounded in dorsal view; width gradually narrowing posteriorly ( Fig. 4D). Occipital carina complete. Malar space 0.5–0.6 (HT: 0.5) × as long as basal width of mandible. Mandible flat at base; lower tooth longer than upper tooth. Antenna with 24 flagellomeres; not flattened and tapped. FL I 2.25–2.5 (HT: 2.5) × as long as maximum depth in lateral view, 1.15–1.25 (HT: 1.25) × as long as FL II.

Mesosoma. Pronotum longitudinally rugulose. Epomia absent. Mesoscutum densely punctate; punctures partly united into groove-like longitudinal foveola ( Fig. 4F); with short notaulus ( Fig. 4D). Scutellum sparsely punctate; flat in lateral view. Mesopleuron longitudinally rugulose dorsally and reticulate rugose ventrally; with conspicuous smooth area around speculum ( Fig. 4E). Epicnemial carina present laterally and ventrally. Sternaulus deep in anterior 0.7 of mesopleuron. Metapleuron reticulate rugose; without juxtacoxal carina. Propodeum rugose or rugulose except for area externa and basalis sparsely punctate with smooth ISP; anterior transverse carina present laterally; posterior transverse carina complete, inverted U-shaped ( Fig. 4F); lateromedian longitudinal carina present, usually indistinct in front of posterior transverse carina; lateral longitudinal carina complete; pleural carina complete; area superomedia partly distinct, transversely rugose; apophysis large, wide, with obtuse apex ( Fig. 4F); spiracle round to oval. Fore wing length 4.25–6.0 (HT: 5.7) mm. Areolet as long as wide; width gradually narrowing anteriorly; received vein 2m-cu at near middle ( Fig. 4G). Fore wing vein 1cu-a interstitial to vein M&RS ( Fig. 4B). Nervellus subvertical; intercepted posterior to middle. Hind femur densely punctate; with long setae ( Figs. 4H, 77F); 4.4–4.6 (HT: 4.4) × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma. T I 2.5–2.75 (HT: 2.5) × as long as maximum width; smooth; longitudinally striate subapically; lateromedian carina absent; dorso-lateral carina present but partly weak. T II 0.7–0.78 (HT: 0.7) × as long as maximum width; smooth; finely and sparsely punctate laterally. Thyridium present; close to anterior margin of T II; slightly depressed; ca. 2.0 × as wide as length. T III to T V finely punctate. Ovipositor sheath 1.0–1.1 (HT: 1.05) × as long as hind tibia, 1.5–1.6 (HT: 1.5) × as long as T I. Ovipositor straight; apex sharp; apex of lower valve with teeth ( Fig. 78C).

Colouration ( Figs. 4A–H). Body (excluding wings) black to blackish-brown. Setae silver. Mandible except for teeth, clypeus, facial orbit, frontal orbit, anterior margin of collar, subtegular ridge, scutellum, postscutellum, posterior margins of T I to T III, and ovipositor reddish-brown to yellowish-brown. Subbasal part of mandible, ventral surfaces of scape and pedicel, palpi, tegula, apex of scutellum, membranous part of metasomal sternites, and apex of metasoma ivory to white. FL V (or VI) to FL XII (or XIII) with white to ivory markings. Legs reddish-brown to brown except for fore and mid coxae, all trochanters, and all trochantelli ivory. Wings hyaline. Veins and pterostigma blackish-brown to brown except for yellowish-brown to ivory wing base.

Male. Unknown.

Distribution. Japan (Amamioshima Is. and Tokunoshima Is.).

Bionomics. Unknown.All species collected in the moist evergreen forest.

Etymology. The specific name is from the Latin “ longi -” (long) plus “ setosus ” (covered with setae), referring to the long setae on the legs.

Remarks. This species resembles Ac. ruficoxalis Sheng & Sun, 2008 in the largely reddish coxae and hind leg and the yellowish face and scutellum but can be distinguished by the hind femur with long setae (with short setae in Ac. ruficoxalis ), the T I 2.5–2.75 × as long as maximum width (2.2 × as long as maximum width in Ac. ruficoxalis ), and the metasoma largely black (largely reddish-brown in Ac. ruficoxalis ). This species also resembles Ac. fujiei sp. nov. in the long setae of hind femur but can be distinguished by the body colouration etc. (see above key).