Cubocephalus asiaticus, Watanabe, 2025

Cubocephalus asiaticus sp. nov.

[New SJN: Asia-onaga-togari-himebachi]

( Figs. 16A–G, 76M, 78M)

Type series. Holotype: JAPAN, KPM-NK 91384 , F, Honshu, Nagano Pref., Ueda City, Sugadaira-kogen , 13– 26. VII. 2015, S. Shimizu leg. (MsT) . Paratype: JAPAN: [Honshu] KPM-NK 102955 , F, Saitama Pref., Naguri Vil., Shomaru-toge , 10–13. X. 1995, A. Shimizu leg. ; KPM-NK 5004434 , F, Kanagawa Pref., Yamakita Vil., Mt. Oomuro-yama , 24. VI. 2013, T. Taniwaki leg. ; KPM-NK 102958 , F, Nagano Pref., Ina City, Hase, Karei-kogen , 30. VII. 2013, S. Yoshizawa leg. ; KPM-NK 102956 , F, Niigata Pref., Myokou City, Suginosawa, Mt. Sasagamine , 20. IX. 2013, S. Shimizu leg. ; KPM-NK 102959 , F, Toyama Pref., Nanto City, Togamura-kamimomose , 21–28. VII. 2009, M. Watanabe leg. (MsT) ; KPM-NK 102960 , F, ditto, 28. VII. – 4. VIII. 2009 ; OMNH, F, ditto, 4–11. VIII. 2009 ; KPM-NK 102961 , F, Toyama Pref., Toyama City, Arimine, Inonedani , 21–28. VII. 2009, M. Watanabe leg. (MsT) ; KPM-NK 102962 , F, ditto, 25. VIII. – 1. IX. 2009 ; KPM-NK 102963 , F, Toyama Pref., Toyama City, Arimine, Jyurodani , 15–22. IX. 2009, M. Watanabe leg. (MsT) ; KPM-NK 102957 , F, Fukui Pref., Ohno City, Kamiuchinami , 29. IX. 1982, T. Murota leg.

Description. Female (n = 12). Body polished; covered with setae; body length 8.9–9.8 (HT: 9.1) mm.

Head 0.6–0.65 (HT: 0.6) × as long as wide in dorsal view. Clypeus 2.5–2.6 (HT: 2.5) × as wide as long; slightly convex in lateral view; punctate dorsally, smooth ventrally; lower margin rounded in frontal view, obtuse in lateral view. Face 0.38–0.4 (HT: 0.38) × as long as minimum width; slightly convex medially; matt with punctures. Frons slightly concave above antennal sockets; coriaceous; punctate dorsally. POL 1.5–2.0 (HT: 2.0) × as OD. OOL 1.4–1.6 (HT: 1.6) × as OD. Gena and occiput finely punctate. Dorsal profile of gena rounded in dorsal view; width not narrowing anteriorly and somewhat abruptly narrowing posteriorly ( Fig. 16D). Occipital carina complete. Malar space 1.0–1.1 (HT: 1.1) × as long as basal width of mandible. Mandible flat at base; lower tooth equal in length of upper tooth. Antenna with 23–25 (HT: 25) flagellomeres; not flattened and tapped. FL I 3.0–3.35 (HT: 3.35) × as long as maximum depth in lateral view, 0.9–0.95 (HT: 0.93) × as long as FL II.

Mesosoma. Pronotum rugose ventrally; densely punctate dorsally ( Fig. 16E). Epomia short; dorsal end situated on collar. Mesoscutum punctate; punctures on median part denser than outer parts. Notaulus sharp ( Fig. 16D); posterior end not reaching centre of mesoscutum. Scutellum punctate; flat in lateral view. Mesopleuron densely punctate and foveolate anteriorly; with conspicuous smooth area around speculum; sparsely punctate postero-ventrally ( Fig. 16E). Epicnemial carina present laterally and ventrally. Sternaulus deep in anterior 0.7 of mesopleuron. Posterior transverse carina of mesosternum without deep notch and teeth. Metapleuron reticulate coriaceous anteriorly, rugulose posteriorly; with complete juxtacoxal carina. Propodeum largely irregularly rugose; anterior transverse carina absent; posterior transverse carina complete, inverted U-shaped ( Fig. 16G); lateromedian longitudinal carina largely indistinct and trace-like; lateral longitudinal carina partly present anteriorly; pleural carina complete; area superomedia indistinct; apophysis absent; spiracle oval. Fore wing length 7.0–7.75 (HT: 7.1) mm. Areolet as long as maximum width; longer than half length of vein 2m-cu; width steeply narrowing anteriorly; received vein 2m-cu at near middle; anterior width shorter than half length of vein 2m-cu ( Fig. 16F). Fore wing vein 1cu-a interstitial to vein M&RS ( Fig. 16F). Nervellus subvertical; intercepted near posterior end of vein. Hind femur reticulate coriaceous; 3.8–4.2 × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma matt and coriaceous. T I 1.65–1.75 (HT: 1.75) × as long as maximum width; latero-median carina absent; dorso-lateral carina complete. Spiracle of T I behind the mid-length of T I. T II 0.7–0.8 (HT: 0.7) × as long as maximum width. Thyridium present; close to anterior margin of T II; flat to slightly depressed; ca. 2.0 × as wide as length. Ovipositor sheath 2.3–2.45 (HT: 2.45) × as long as hind tibia, 3.85–4.25 (HT: 3.85) × as long as T I. Ovipositor weakly upcurved; apex sharp and without minute teeth dorsally; apex of lower valve with teeth ( Fig. 78M).

Colouration ( Figs. 16A–G). Body (excluding wings) black to blackish-brown. Setae silver. Subapical part of mandible tinged with reddish-brown. Labrum reddish-brown. FL VI (or VII) to FL X with white markings. Bases of femora, all tibial spurs, and fore tibia more or less tinged with brown to reddish-brown. Posterior margins of metasomal tergites sometimes narrowly tinged with reddish-brown. Postero-median membranous parts of T V to T VII white. Thyridium and ovipositor reddish-brown. Wings hyaline. Veins and pterostigma blackish-brown to brown except for yellowish wing base.

Male. Unknown.

Distribution. Japan (Honshu).

Bionomics. Unknown.

Etymology. The specific name is from Asia, referring to the distribution of this species, Japan, is a part of Asia.

Remarks. This species resembles Cu. atrator and Cu. confusus sp. nov. in the entirely black legs and almost or entirely black metasomal tergites but can be distinguished by the posterior transverse carina of mesosternum without deep notch and teeth (with deep notch and teeth in Cu. atrator ), FL I 3.0–3.35 × as long as maximum depth in lateral view (3.9 × in Cu. atrator ; 2.1–2.4 × in Cu. confusus ) and the ovipositor sheath 2.3–2.45 × as long as hind tibia (1.83–2.68 × in Cu. atrator ; 1.8–2.0 × in Cu. confusus ).