Cubocephalus sapporensis, Watanabe, 2025

Cubocephalus sapporensis sp. nov.

[New SJN: Sapporo-onaga-togari-himebachi]

( Figs. 21A–G, 76P, 78Q)

Type series. Holotype: JAPAN, KPM-NK 91382 , F, Hokkaido, Sapporo City, Mt. Maruyama , 29. VII. 2009, K. Watanabe leg.

Description. Female (n = 1). Body matt; covered with setae; body length 12.4 mm.

Head 0.63 × as long as wide in dorsal view. Clypeus 2.0 × as wide as long; slightly convex in lateral view; coriaceous and densely punctate dorsally, sparsely punctate with some transverse foveae ventrally; lower margin subtruncate in frontal view, obtuse in lateral view. Face 0.4 × as long as minimum width; weakly convex and densely punctate medially. Frons slightly concave above antennal sockets; densely punctate except for smooth areas above antennal sockets. POL 1.5 × as OD. OOL 1.5 × as OD. Gena and occiput finely and densely punctate. Dorsal profile of gena rounded in dorsal view; width gradually narrowing posteriorly ( Fig. 21D). Occipital carina complete. Malar space 1.2 × as long as basal width of mandible. Mandible flat at base; lower tooth equal in length of upper tooth. Antenna with 25 flagellomeres; not flattened and tapped. FL I 3.15 × as long as maximum depth in lateral view, 1.1 × as long as FL II.

Mesosoma. Pronotum rugose ventrally; rugulose dorsally. Epomia short; dorsal end situated slightly above collar. Mesoscutum densely punctate. Notaulus sharp; posterior end not reaching centre of mesoscutum. Scutellum punctate; flat in lateral view. Mesopleuron finely and irregularly rugulose; with conspicuous smooth area around speculum ( Fig. 21E). Epicnemial carina present laterally and ventrally. Sternaulus deep in entire length of mesopleuron. Posterior transverse carina of mesosternum without deep notch and teeth. Metapleuron reticulate coriaceous anteriorly, rugulose posteriorly; with complete (but weak medially) juxtacoxal carina. Propodeum rugulose to rugose; anterior transverse carina slightly present as trace-like; posterior transverse carina complete, inverted V-shaped ( Fig. 21G); lateromedian longitudinal carina complete but entirely weak; lateral longitudinal carina complete; pleural carina complete; area superomedia partly defined, longer than wide; apophysis absent; spiracle oval. Fore wing length 8.5 mm. Areolet as long as maximum width; as long as half-length of vein 2m-cu; width steeply narrowing anteriorly; received vein 2m-cu at near middle; anterior width shorter than half length of vein 2m-cu ( Fig. 21F). Fore wing vein 1cu-a interstitial to vein M&RS. Nervellus subvertical; intercepted near posterior end of vein. Hind femur reticulate coriaceous; 4.6 × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma. T I 2.2 × as long as maximum width; lateromedian carina absent; dorso-lateral carina absent. Spiracle of T I behind the mid-length of T I; base convex ( Fig. 21G). T II 0.85 × as long as maximum width. Thyridium present; close to anterior margin of T II; slightly depressed; ca. 2.0 × as wide as length. Ovipositor sheath 1.65 × as long as hind tibia, 3.15 × as long as T I. Ovipositor weakly upcurved; apex sharp and without minute teeth dorsally; apex of lower valve with many (more than 16) teeth ( Fig. 78Q).

Colouration ( Figs. 21A–G). Body (excluding wings) black to blackish-brown. Setae silver. Subapical part of mandible, lateral sides of clypeus, ventral surface of apical part of flagellum, and postero-dorsal corner of pronotum tinged with reddish-brown. FL V to FL XI with white markings. T II and T III largely tinged with dark reddish-brown. Membranous part of metasomal sternites yellowish-brown. Trochanters, trochantelli, bases of femora, fore femur, fore tibia, fore and mid femora, and fore tibial spurs partly tinged with brown. Median membranous part of T VII white. Thyridium and ovipositor reddish-brown. Wings hyaline. Veins and pterostigma blackish-brown to brown except for yellowish wing base.

Distribution. Japan (Hokkaido).

Bionomics. Unknown.

Etymology. The species name is from the type locality, Sapporo, a city in Hokkaido.

Remarks. This species resembles Cu. atrator and C. uryuensis sp. nov. in the largely black body colouration but can be distinguished by the short ovipositor sheath (see above key).