Javra japonica, Watanabe, 2025

Javra japonica sp. nov.

[New SJN: Nippon-kuro-togari-himebachi]

( Figs. 32A–K, 79C)

Javra sp. A & E: Watanabe & Taniwaki, 2018: 80.

Type series. Holotype: JAPAN, KPM-NK 102795 , F, Honshu, Toyama Pref., Toyama City, Arimine , Jyurodani , 21–28. VII. 2009, M. Watanabe leg. (MsT) . Paratype: JAPAN [Honshu] KPM-NK 102813 , F, Saitama Pref., Tokigawa Town, Oono-toge , 15. X. 2006, T. Mita leg.; KPM-NK 102814 , F, Nagano Pref., Ina City, Hase, Karei-kogen , 30. VII. 2013, S. Yoshizawa leg.; OMNH ,

1F, Toyama Pref., Nanto City, Togamura-kamimomose , 11–16. VIII. 2009, M. Watanabe leg. (MsT); KPM-NK 102812 , ditto, 11–18. VIII. 2009; KPM-NK 102796 , F, Toyama Pref., Toyama City, Arimine, Jyurodani , 11–16. VIII. 2009, M. Watanabe leg. (MsT); KPM-NK 102797 , F, ditto, 16–25. VIII. 2009; KPM-NK , F, 102810, ditto, 25. VIII. – 1. IX. 2009; KPM-NK 102811 , F, ditto, 15–22. IX. 2009; KPM-NK 102817 , F, Toyama Pref., Toyama City, Arimine, Inone-dani , 7–14. VII. 2009, M. Watanabe leg. (MsT); KPM-NK 102816 , F, Toyama Pref., Toyama City, Kamegai , 28. VII. – 4. VIII. 2009, M. Watanabe leg. (MsT); KPM-NK 102818 , F, Fukui Pref., Ikeda Town, Kouchi , 17. VI. 1981, H. Kurokawa leg.

Description. Female (n = 12). Body polished; covered with setae; body length 6.0–8.4 (HT: 7.1) mm.

Head 0.6–0.65 (HT: 0.62) × as long as wide in dorsal view. Clypeus 1.8–1.9 (HT: 1.9) × as wide as long; slightly convex in lateral view; sparsely punctate; lower margin subtruncate; sharp in lateral view. Face 0.5 × as long as minimum width; matt; slightly convex medially. Anterior tentorial pit small. Frons slightly concave above antennal sockets; coriaceous; sparsely punctate except for area above antennal sockets. POL 0.9–1.0 (HT: 0.9) × as OD. OOL 1.0–1.1 (HT: 1.0) × as OD. Dorsal profile of gena straight in dorsal view; width abruptly narrowing posteriorly ( Fig. 32D). Occipital carina complete. Malar space 1.2–1.3 (HT: 1.2) × as long as basal width of mandible. Mandible flat at base; lower tooth equal in length of upper tooth. Antenna with 26–28 (HT: 27) flagellomeres; not flattened and tapped. FL I 4.35–5.0 (HT: 5.0) × as long as maximum depth in lateral view, 1.15–1.25 (HT: 1.25) × as long as FL II.

Mesosoma. Pronotum rugulose ventrally; sparsely punctate dorsally. Epomia short; dorsal end situated slightly above collar. Mesoscutum densely punctate. Notaulus sharp; posterior end reaching centre of mesoscutum or slightly shorter. Scutellum densely punctate; weakly convex in lateral view. Mesopleuron longitudinally rugulose dorsally, irregularly rugulose ventrally; with conspicuous small smooth area around speculum ( Fig. 32E). Epicnemial carina present laterally and ventrally. Sternaulus deep in entire length of mesopleuron. Metapleuron largely punctate; with partly indistinct or complete juxtacoxal carina. Propodeum punctate anteriorly; rugose or rugulose posteriorly ( Figs. 32H–J); with all carinae except for anterior transverse carina largely indistinct; lateral section of anterior transverse carina weak and not clearly defined by transverse rugae on area dentipara; area superomedia defined, slightly longer than maximum width; area superomedia and dentipara largely covered with transverse rugae; apophysis short; apex obtuse; spiracle oval. Fore wing length 5.1–7.0 (HT: 6.35) mm. Areolet as long as maximum width or sometimes wider than long; width steeply narrowing anteriorly; received vein 2m-cu at near middle ( Figs. 32F, G). Fore wing vein 1cu-a interstitial to vein M&RS. Nervellus subvertical; intercepted near posterior end of vein. Hind femur reticulate coriaceous; 5.1–5.5 (HT: 5.2) × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma coriaceous ( Fig. 32K). T I 2.45–2.5 (HT: 2.45) × as long as maximum width; latero-median carina obtusely present except for apical part; dorso-lateral carina complete. T II 0.95–1.0 (HT: 0.95) × as long as maximum width. Thyridium present; somewhat distant from (by more than length of thyridium) anterior margin of T II; flat to slightly depressed; ca. 2.0 × as wide as length. Ovipositor sheath 0.88–1.03 (HT: 0.95) × as long as hind tibia, 1.6– 2.05 (HT: 1.9) × as long as T I. Ovipositor straight; apex sharp; apex of lower valve with teeth ( Fig. 79C).

Colouration ( Figs. 32A–K). Body (excluding wings) black to blackish-brown. Setae silver. Mandible except for teeth, clypeus, and posterior margins of each metasomal tergite tinged with reddish-brown. Ventral surface of scape, pedicel, and apical part of flagellum and tegula sometimes tinged with brown to reddish-brown. FL VI to FL IX with white markings. Scutellum and postscutellum usually each with ivory to yellow marking; marking sometimes changed as reddish-brown marking or reduced. Trochanters, trochantelli, and base of femora sometimes narrowly tinged with yellow. Fore and mid legs except for coxa usually partly tinged with brown. Base of hind tibia brown to ivory. Membranous part of metasomal sternites yellow to yellowish-brown. Thyridium and ovipositor reddish-brown. Wings hyaline. Veins and pterostigma blackish-brown to brown except for yellowish-brown to yellow wing base.

Male. Unknown.

Distribution. Japan (Honshu).

Bionomics. Unknown.

Etymology. The specific name is from Japan.

Remarks. This species has the scutellum with or without ivory to yellow marking. While I found intermediate condition of both character states, reddish-brown coloured marking on scutellum in KPM-NK 102796. I also found no morphological difference between them and thus, I conclude that it is intraspecific variation of this species. This species resembles J. jemilleri ( Kriechbaumer, 1893) in the body colouration and the length of ovipositor sheath but can be distinguished the anterior transverse carina of propodeum largely absent (present its lateral section in J. jemilleri ) and the mesopleuron largely rugulose (largely granulate in J. jemilleri ).