Javra gigantea, Watanabe, 2025

Javra gigantea sp. nov.

[New SJN: Katayama-togari-himebachi]

( Figs. 31A–H, 79B)

Type series. Holotype: JAPAN, KPM-NK 102819 , F, Honshu, Tochigi Pref., Nasushiobara City, Komakigawarindo , 7. IX. 2021, E. Katayama leg. Paratype: JAPAN, KPM-NK 102820 , F, Honshu, Fukui Pref., Ohno City, Dosai-zan , 23. V. 1982, T. Murota leg.

Description. Female (n =2). Body matt; covered with setae; body length 9.25–11.3 (HT: 11.3) mm.

Head 0.6 × as long as wide in dorsal view. Clypeus 2.1–2.2 (HT: 2.1) × as wide as long; slightly convex in lateral view; smooth; sparsely punctate; lower margin subtruncate; sharp in lateral view. Face 0.4 × as long as minimum width; slightly convex medially. Anterior tentorial pit small. Frons slightly concave above antennal sockets; transversely rugulose above antennal sockets; with pair of slight convexities just above mid-height ( Fig. 31E). POL 1.3–1.35 (HT: 1.3) × as OD. OOL 1.3–1.5 (HT: 1.3) × as OD. Occiput and vertex finely and densely punctate. Dorsal profile of gena weakly rounded in dorsal view; width gradually narrowing posteriorly ( Fig. 31D). Occipital carina complete. Malar space 1.1–1.2 (HT: 1.2) × as long as basal width of mandible. Mandible flat at base; lower tooth equal in length of upper tooth. Antenna with 26–28 (HT: 28) flagellomeres; not flattened and tapped. FL I 5.25 × as long as maximum depth in lateral view, 1.05 × as long as FL II.

Mesosoma. Pronotum rugulose and rugose. Epomia short; dorsal end situated slightly above collar. Mesoscutum densely punctate. Notaulus sharp; posterior end reaching centre of mesoscutum. Scutellum punctate; weakly convex in lateral view. Mesopleuron rugulose ( Fig. 31F); with conspicuous smooth area around speculum. Epicnemial carina present laterally and ventrally. Sternaulus deep in entire length of mesopleuron. Metapleuron rugose; with complete juxtacoxal carina. Propodeum rugose except for area basalis and area externa coriaceous; rugose or rugulose posteriorly; with all carinae ( Fig. 31H); lateral longitudinal carina partly indistinct; area superomedia defined, slightly longer than maximum width; median part of posterior transverse carina higher than other parts and strongly raised; area postero and area dentipara largely covered with oblique rugae; apophysis short; apex obtuse; spiracle oval. Fore wing length 7.4–9.8 (HT: 9.8) mm. Areolet longer than maximum width; width gradually narrowing anteriorly; received vein 2m-cu at near middle ( Fig. 31G). Fore wing vein 1cu-a slightly antefurcal to vein M&RS ( Fig. 31G). Nervellus subvertical; intercepted near posterior end of vein. Hind femur reticulate coriaceous; 6.0–6.2 (HT: 6.2) × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma. T I 2.0–2.2 (HT: 2.2) × as long as maximum width; latero-median carina indistinct; dorso-lateral carina absent or partly present posteriorly. T II 0.75 × as long as maximum width; thyridium indistinct. Ovipositor sheath 1.18–1.23 (HT: 1.23) × as long as hind tibia, 2.4–2.5 (HT: 2.4) × as long as T I. Ovipositor straight; apex sharp; apex of lower valve with teeth ( Fig. 79B).

Colouration ( Figs. 31A–H). Body (excluding wings) black to blackish-brown. Setae silver. Subapical part of mandible weakly tinged with reddish-brown. Face with pair of yellow markings between antennal sockets and eye. FL VI (or VII) to FL IX with white markings. Hind second to fourth tarsomeres ivory. Ovipositor reddish-brown. Wings hyaline. Veins and pterostigma blackish-brown to brown except for yellowish-brown wing base.

Male. Unknown.

Distribution. Japan (Honshu).

Bionomics. Unknown.

Etymology. The specific name is from the Latin “ giganteus ” (giant), referring to the largest body size in Japanese species.

Remarks. This species resembles J. tenuis sp. nov. in the large body with white banded hind tibia but can be distinguished by the shape of areolet and the surface of mesoscutum (see above key). This species also resembles J. coreensis and J. taniguchiae in the black body but can be distinguished by the large body size (more than 9.0 mm in this species; less than 8.0 mm in J. coreensis and J. taniguchiae ) and the hind tibia without white base (with white base in J. taniguchiae ).