Javra albotrochantellata, Watanabe, 2025

Javra albotrochantellata sp. nov.

[New SJN: Ashi-shiromon-togari-himebachi]

( Figs. 30A–H, 79A)

Aptesis sp. D : Watanabe & Taniwaki, 2018: 75.

Javra sp. B : Watanabe & Taniwaki, 2018: 80.

Type series. Holotype: JAPAN, KPM-NK 102847 , F, Honshu, Saitama Pref., Hanno City, Naguri Vil., Shomaru-toge , 10–13. X. 1995, A. Shimizu leg. Paratype: JAPAN: [Hokkaido] OMNH, KPM-NK 102823 , and 102850, 3F, Horokanai Town, Uryu , 11–17. VII. 2012, K. Watanabe leg. (MsT) . [ Honshu ] KPM-NK 102848 , F, Yamanashi Pref., Koushu City, Sagashio , 16. VI. 2007, K. Watanabe leg. ; KPM-NK 102849 , F, Nagano Pref., Neba Vil., Chayagawa , 19. IX. 2014, M. Takakuwa leg. ; KPM-NK 102822 , F, Toyama Pref., Toyama City, Kamegai , 28. VII. – 4. VIII. 2009, M. Watanabe leg. (MsT).

Description. Female (n = 7). Body polished; covered with setae; body length 5.0–9.8 (HT: 9.8) mm.

Head 0.58–0.6 (HT: 0.58) × as long as wide in dorsal view. Clypeus 1.8–2.0 (HT: 1.9) × as wide as long; slightly convex in lateral view; sparsely punctate; lower margin subtruncate or slightly concave medially; with transverse shallow groove along lower margin; sharp in lateral view. Face 0.4 × as long as minimum width; matt; finely punctate medially; slightly convex medially. Anterior tentorial pit small. Frons slightly concave above antennal sockets; coriaceous; sparsely punctate dorsally; area above antennal sockets smooth. POL 0.9–1.0 (HT: 0.95) × as OD. OOL 1.0–1.5 (HT: 1.1) × as OD. Occiput and vertex finely and densely punctate. Dorsal profile of gena straight in dorsal view; width gradually narrowing posteriorly ( Fig. 30D). Occipital carina complete. Malar space 1.0–1.2 (HT: 1.2) × as long as basal width of mandible. Mandible flat at base; lower tooth equal in length of upper tooth. Antenna with 23–27 (HT: 27) flagellomeres; not flattened and tapped. FL I 3.65–4.0 (HT: 3.65) × as long as maximum depth in lateral view, 1.0–1.1 (HT: 1.0) × as long as FL II.

Mesosoma. Pronotum rugulose ventrally; finely and densely punctate dorsally. Epomia short; dorsal end situated slightly above collar. Mesoscutum densely punctate. Notaulus sharp; posterior end not reaching centre of mesoscutum. Scutellum densely punctate; weakly convex in lateral view. Mesopleuron finely punctate; largely longitudinally rugulose; with conspicuous smooth area around speculum ( Fig. 30E). Epicnemial carina present laterally and ventrally. Sternaulus deep in anterior 0.6 of mesopleuron. Metapleuron densely punctate anteriorly, coriaceous posteriorly; with complete juxtacoxal carina. Propodeum coriaceous ( Fig. 30G); with all carinae but anterior transverse carina, lateromedian longitudinal carina, and lateral longitudinal carina weak and sometimes partly indistinct (anterior transverse carina sometimes largely absent in small specimens); area superomedia at least partly defined, longer than maximum width; apophysis short; apex obtuse; spiracle oval. Fore wing length 4.1–7.8 (HT: 7.8) mm. Areolet as long as maximum width; width gradually to steeply narrowing anteriorly; received vein 2m-cu at near middle ( Fig. 30F). Fore wing vein 1cu-a interstitial to vein M&RS ( Fig. 30F). Nervellus subvertical; intercepted near posterior end of vein. Hind femur reticulate coriaceous; 4.6–5.1 (HT: 5.1) × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma. T I 1.95–2.4 (HT: 2.35) × as long as maximum width; largely matt ( Fig. 30H); latero-median carina obtusely present except for apical part; dorso-lateral carina complete. T II 0.85–0.95 (HT: 0.95) × as long as maximum width; matt. Thyridium present; somewhat distant from (by more than length of thyridium) anterior margin of T II ( Fig. 30H); flat to slightly depressed; ca. 1.0 × as wide as length. T III to T V slightly coriaceous to smooth; finely punctate. Ovipositor sheath 0.85–1.1 (HT: 0.85) × as long as hind tibia, 1.6–2.25 (HT: 1.6) × as long as T I. Ovipositor straight; apex sharp; apex of lower valve with teeth ( Fig. 79A).

Colouration ( Figs. 30A–H). Body (excluding wings) black to blackish-brown. Setae silver. Subapical part of mandible and posterior margins of each metasomal tergite tinged with reddish-brown. FL VI to FL X (or IX) with white markings. Trochanters, trochantelli, and apex of metasoma ivory; sometimes fore trochanter weakly darkened. Membranous part of metasomal sternites dark yellowish-brown. Thyridium and ovipositor reddish-brown. Wings hyaline. Veins and pterostigma blackish-brown to brown except for yellowish-brown to yellow wing base.

Male. Unknown.

Distribution. Japan (Hokkaido and Honshu).

Bionomics. Unknown.

Etymology. The specific name is from Latin “ albi ” (white) plus “ trochantellata ” (trochantellus), referring to the white trochantelli of all legs.

Remarks. This species resembles J. coreensis and J. taniguchiae in the black body but can be distinguished by the whitish hind trochanter and trochantellus (black in J. coreensis and J. taniguchiae ) and the ovipositor sheath 0.85–1.1 × as long as hind tibia (0.65–0.8 × in J. coreensis ; 1.43 × in J. taniguchiae ) (see above key).