Parmortha gigantea, Watanabe, 2025

Parmortha gigantea sp. nov.

[New SJN: Ito-onaga-togari-himebachi]

( Figs. 45A–G, 77C, J, 79N)

Type series. Holotype: JAPAN, KPM-NK 75808 , F, Hokkaido, Sapporo City, Teinekanayama , 18. VII. 2012, M. Ito leg. Paratype: JAPAN, KPM-NK 103049 , F, Honshu, Toyama Pref., Nanto City, Togamura-kamimomose , 28. VII. – 4. VIII. 2009, M. Watanabe leg. (MsT) .

Description. Female (n= 2). Body matt; lustre dull; covered with setae; body length 10.5–13.9 (HT: 13.9) mm.

Head 0.6 × as long as wide in dorsal view. Clypeus 2.1 × as wide as long; slightly convex in lateral view; punctate dorsally, sparsely punctate ventrally, ISP smooth; lower margin subtruncate in frontal view, obtuse in lateral view. Face 0.45 × as long as minimum width; slightly convex medially ( Fig. 45C); finely and densely punctate. Frons slightly concave above antennal sockets; finely and densely punctate except for smooth areas above antennal sockets. POL 1.0–1.15 (HT: 1.0) × as OD. OOL 1.3–1.5 (HT: 1.3) × as OD. Gena and occiput finely punctate. Dorsal profile of gena rounded in dorsal view; width not narrowing anteriorly and somewhat abruptly narrowing posteriorly ( Fig. 45D). Occipital carina complete. Malar space 1.0–1.05 (HT: 1.0) × as long as basal width of mandible. Mandible flat at base; lower tooth equal in length of upper tooth. Antenna with 29 flagellomeres; not flattened and tapped. FL I 5.0–5.7 (HT: 5.0) × as long as maximum depth in lateral view, 1.0 × as long as FL II.

Mesosoma. Pronotum striate ventrally, finely and densely punctate dorsally ( Fig. 45E). Epomia short; section on border of collar and pronotum present. Mesoscutum densely and finely punctate, ISP largely smooth. Notaulus long and sharp; posterior end close to centre of mesoscutum. Scutellum punctate; weakly convex in lateral view. Mesopleuron with conspicuous smooth area around speculum; obliquely or longitudinally striate ( Fig. 45E). Epicnemial carina present laterally and ventrally. Sternaulus deep in entire length of mesopleuron. Metapleuron with complete, weak juxtacoxal carina. Propodeum irregularly rugae and reticulate rugulose; anterior transverse carina present as trace-like by rugae; posterior transverse carina complete ( Fig. 45G); lateromedian longitudinal carina as trace-like by rugae; lateral longitudinal carina present, weak anteriorly; pleural carina complete; area superomedia defined but partly indistinct; apophysis absent; spiracle oval. Fore wing length 8.5–10.8 (HT: 10.8) mm. Areolet slightly shorter than maximum width; longer than half length of vein 2m-cu; width gradually narrowing anteriorly; received vein 2m-cu basal than middle; anterior width longer than half length of vein 2m-cu ( Fig. 45F). Fore wing vein 1cu-a interstitial to vein M&RS ( Fig. 45F). Nervellus subvertical; intercepted posterior to middle ( Fig. 45F). Hind femur reticulate coriaceous; 5.2–5.4 (HT: 5.2) × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma. T I 1.9–2.0 (HT: 2.0) × as long as maximum width; latero-median carina absent; dorso-lateral carina partly and weakly present. Spiracle of T I situated near the mid-length of T I ( Fig. 77J). T II 0.75–0.9 (HT: 0.9) × as long as maximum width. Thyridium present; close to anterior margin of T II; flat; ca. 2.0 × as wide as length. Ovipositor sheath 2.6–2.8 (HT: 2.8) × as long as hind tibia, 5.9–6.4 (HT: 6.4) × as long as T I. Ovipositor upcurved; apex sharp and with some minute teeth dorsally; apex of lower valve with teeth ( Fig. 79N).

Colouration ( Figs. 45A–G). Body (excluding wings) black to blackish-brown. Setae silver. FL VI to FL IX (or X) with white markings. Fore and mid tibiae and tarsi partly tinged with dark brown. Apical half of hind TS II, hind TS III, and hind TS IV ivory to white. Posterior margins of T V and T VI each narrowly tinged with ivory to white. Postero-median membranous part of T VII ivory to white. Ovipositor reddish-brown. Wings hyaline. Veins and pterostigma blackish-brown to brown.

Male. Unknown.

Distribution. Japan (Hokkaido and Honshu).

Bionomics. Unknown.

Etymology. The specific name is from the Latin “ giganteus ” (giant), referring to the largest body size in Japanese species.

Remarks. This species resembles Pa. nigra sp. nov. (see remarks of Pa. nigra ). This species also resembles Echthrus species in the spiracle of T I but can be easily distinguished by the normal shaped fore tibia ( Fig. 77C) (fore tibia significantly enlarged in Echthrus ).