Goryphus albofasciatus ( Matsumura & Uchida, 1926 )

Goryphus albofasciatus ( Matsumura & Uchida, 1926) View in CoL

[SJN: Shiro-yokojima-togari-himebachi]

( Figs. 64A–J, 65A–F, 80F)

Gambrus albofasciatus Matsumura & Uchida, 1926: 75 .

Description. Female (n = 10). Body polished; covered with setae; body length 7.3–11.2 mm.

Head 0.6–0.65 × as long as wide in dorsal view. Clypeus 1.8–2.0 × as wide as long; weakly convex in lateral view; densely punctate dorsally; smooth ventrally; lower margin rounded in frontal view, narrowly reflected in lateral view. Face 0.5–0.55 × as long as minimum width; slightly convex medially; punctate laterally; foveola rugose medially. Anterior tentorial pit small. Frons weakly concave above antennal sockets; rugose dorsally except for coriaceous lateral sides; smooth on concavity. POL 0.6–1.0 × as OD. OOL 1.0–1.3 × as OD. Gena and occiput finely and densely punctate, ISP coriaceous. Dorsal profile of gena slightly rounded to straight in dorsal view; width abruptly narrowing posteriorly. Occipital carina complete. Malar space 0.7–0.75 × as long as basal width of mandible. Mandible slightly convex at base; lower margin not widened as lamella; lower tooth shorter than upper tooth. Antenna with 25–27 flagellomeres; apical part slightly widened. FL I 8.8–10.5 × as long as maximum depth in lateral view, 1.05–1.1 × as long as FL II.

Mesosoma. Pronotum reticulate rugose. Epomia long; dorsal end closed to dorsal margin of pronotum and strongly raised as convexity. Mesoscutum densely and finely punctate; rugulose posteromedially; with long and strong notaulus. Scutellum punctate; convex in lateral view; with lateral carina on anterior half. Mesopleuron largely irregularly rugose; with smooth area around speculum ( Fig. 64E). Epicnemial carina present laterally and ventrally; dorsal end reaching below subtegular ridge. Sternaulus deep in entire length of mesopleuron. Metapleuron irregularly rugose; with complete juxtacoxal carina. Propodeum rugose ( Fig. 64H): anterior transverse carina complete; posterior transverse carina largely indistinct; lateral longitudinal carina absent; pleural carina present; lateromedian longitudinal carina present only anterior to anterior transverse carina; apophysis large and ear-like ( Figs. 64E, H); spiracle oval. Fore wing length 5.6–8.5 mm. Areolet small; as long as maximum width; width slightly narrowing anteriorly; received vein 2m-cu near middle ( Figs. 64F, G). Fore wing vein 1cu-a slightly antefurcal to vein M&RS ( Figs. 64F, G). Nervellus subvertical; intercepted posterior to middle ( Fig. 64F). Hind 5.2–5.75 × as long as maximum depth in lateral view. Tarsal claws simple.

Metasoma densely punctate except for T I ( Figs. 64I, J). T I 1.55–1.75 × as long as maximum width; sparsely punctate except for smooth basal part; latero-median carina present except for posterior part; dorso-lateral carina complete; with pair of lateral triangular teeth at base. T II 0.9–0.95 × as long as maximum width; thyridium weakly present close to anterior margin of T II; slightly depressed; ca. 2.0 × as wide as length. Ovipositor sheath 0.85–0.9 × as long as hind tibia, 1.7–2.05 × as long as T I. Ovipositor straight; apex sharp; nodus slightly present; apex of lower valve with teeth ( Fig. 80F).

Colouration ( Figs. 64A–J). Body (excluding wings) black to blackish-brown. Setae silver. FL V (or VI) to FL XI with white markings. T II and T VII with white marking(s). Palpi, fore and mid legs, and membranous parts of metasomal sternites sometimes tinged with ivory. Ovipositor reddish-brown. Some tarsal segment(s) of legs ivory. Wings hyaline; with clouded area. Veins and pterostigma blackish-brown except for brown to yellowish-brown wing base. Some intersubspecific variation of coloration present; see description of G. a. erabu ssp. nov.

Male (n =15). Similar to female ( Figs. 65A–F). Body length 6.0– 10.7 mm. Head 0.55 × as long as wide in dorsal view. Face 0.6–0.65 × as long as minimum width. Antenna with 26–31 flagellomeres; with tyloids on FL XIV (or XIII) to FL XV (or XVI). FL I 3.55–3.85 × as long as maximum depth in lateral view, 1.25 × as long as FL II. Fore wing length 4.3–7.3 mm. T I 2.0–2.6 × as long as maximum width. T II 1.0–1.5 × as long as maximum width. Antenna without white band. Mandible, median part of collar, and fore and mid coxae and trochanters with ivory area. T III with white posterior band. Clouded area of fore wing weaker than female. Whitish parts of legs larger than female.

D i s t r i b u t i o n. J a p a n (O k i n a w a j i m a I s. a n d Okinoerabujima Is.).

Bionomics. Unknown. Adults were collected in evergreen forests.

Remarks. This species is easily distinguished from other Japanese species by the black and white body colouration. Matsumura & Uchida (1926) described this species based on the specimens collected from Okinawajima Island. Momoi (1970) recorded additional data of this species based on the specimens collected in Okinawajima Island and Okinoerabujima Island. I found some additional materials of this species from both islands and recognized stable differences of body colouration. In this study, I newly describe a new subspecies of this species based on the specimens collected from Okinoerabujima Is.