Calomera paradecemguttata, Moravec & Dheurle & Schüle & Wiesner, 2025

Calomera paradecemguttata sp. nov.

( Figs 48–98 View FIGURES 48–51 View FIGURES 52–55 View FIGURES 56–64 View FIGURES 65–74 View FIGURES 75–92 View FIGURES 93–98 )

Cicindela decemguttata sensu auctorum, partim. Non C. decemguttata Fabricius, 1801 View in CoL !

Type locality. Indonesia: S.O. Sulawesi, Wakarumba, Buton Island .

Type material. Holotype ♂ in SDEI, labelled: “ Indonesia, Sulawesi S.O. / Ins. Buton - Wakarumba / 3-7.II.1994 / M. Štrba & I. Jeniš leg.” [printed] // “ Holotype / Calomera paradecemguttata sp. nov. / (= C. decemguttata sensu auct.) / det. J. Moravec, C. Dheurle / P. Schüle & J. Wiesner 2024” [red, printed] . Allotype. ♀ in CCJM with same label data as holotype . Paratypes. 5 ♂♂, 1 ♀ in CCJM , 3 ♂♂ in JWCM , 1 ♂ in MHCP with same label data as holotype . 4 ♂♂ in CCJM: “ Sulawesi S. O. / Kendari airport / 11-14.II.1994 / M. Štrba & I. Jeniš lgt.” . 1 ♂, 3 ♀♀ in NHMW: “S.E-Sulawesi / Kendari airport 11-14.2. / 30km W of Kendari 1994 / leg, M. Strba & I. Jenis lgt.” . 2 ♂♂, 1 ♀ in NHMW: “Z Sulawesi, 29.1.1995 / Salope, Wasserfall / leg. R. Seyfert ”. 1 ♂ in NHMW: “Z. Sulawesi, Boe / S Ufer Poso See / 15.4.1995 / leg. R. Gerstmeier ” // “Collection J. Probst”. 3 ♀♀ in NHMW: “S. Sulawesi / Mt. Lompo Batang / March 1992 ” // “ Lophyridia (s. str.) / decemguttata / ( Fabricius, 1801) / det. J. Probst ”. 6 ♂♂, 3 ♀♀ in NHMW, 3 ♂♂, 1 ♀ in CCJM (ex NHMW): “ Indonesia, C-Sulawesi / Lore-Lindu-NP / Kamorora, 13.- 18.4.1994 / leg. R. Gerstmeier ” // “ Lophyridia (s. str.) / decemguttata ( Fabricius 1801) / det. J. Probst ” // “Collection J. Probst”. 1 ♀ in NHMW: “ Sulawesi 1992 / Tasek Ria / W. Manado / leg. Jäch, 13.IV.” // “ Lophyridia (s. str.) / decemguttata ( Fabricius 1801) / det. J. Probst ”. 2 ♂♂ in NHMW, 1 ♀ in SDEI: “ Nord Celebes / Toli-Toli / Nov.-Dec.1895 / H. Fruhstorfer // Coll. Moosbrugger” // “Aqu-Nr. / 1954-74”. 1 ♀ in CDCL: “ Nord-Celebes / Toli - Toli / Nov-Dez 1895” [printed] // “C. 10-guttata / Fab. ” [handwritten]. 1 ♂ in MFNB, 2 ♂♂ in BMNH: “ Nord-Celebes / Toli -Toli / / Nov-Dez 1895 / H. Fruhstorfer ”. 1 ♂ in NMPC: “ Nord Celebes / Toli-Toli ”. 1 ♂ in NMPC: “ Staudinger / Minahassa ” [handwritten] // “ Coll. Srnka ”. 1 ♂ in NHMW: “N. Sulawesi 1992 / Wakan (10) / Amurang – Motoling / leg. Jäch 18.IV.”. 1 ♀ in NHMW: “N. Sulawesi 1992 / Umg. Modoinding / NE Kotamobagu (12) / leg. Jäch 18.IV.”. // “ Lophyridia (s. str.) / decemguttata ( Fabricius 1801) / det. J. Probst ”. 1 ♀ in NHMW: “N. Sulawesi 1992 / Dua Saudara NP / W-Batuputih (8) / leg. Jäch 18.IV.”. // “ Lophyridia (s. str.) / decemguttata ( Fabricius 1801) / det. J. Probst ”. 1 ♂ in NHMW “ Palolo Palu / C. Sulawesi / 14.I.1968 Jasmin leg.”. 1 ♀ in NHMW: “N. Sulawesi 1992 / Bitauna, Kotamo- / bagu-Gorontalo (18) / leg. Jäch 18.IV.”. // “ Lophyridia (s. str.) / decemguttata ( Fabricius 1801) / det. J. Probst ”. 2 ♂♂, 1 ♀ in JWCM: “18.- 19.8.1990, S-Sulawesi, 15 km W Palopo, leg. A. Riedel ”. 1 ♀ in JWCM: “25.- 27.8.1990, C-Sulawesi, Palu, Palolo, Lindu-N.P. , leg. A. Riedel ”. 2 ♂♂, 2 ♀♀ in JWCM: “C Sulawesi / Palolo, Palu / 10.1990”. 1 ♂ in NMPC: “ Indonesia C. Sulawesi, W / Poso, 11-16.iv.1999, 1–400 / m, 5-10km SW Tambarana / Bečvář & Zábranský leg.”. 1 ♂ in NMPC: Sulawesi / Toraja / Palu [Palolu] Palu / I.1999 / Bečvář lgt.”. 3 ♂♂ in NMPC, 1 ♂ in MKPC: “ Sulawesi S / 20 km NE Sabbang / 5.VI.2001 Bolm lgt.” [handwritten]. 1 ♂ in NMPC: “S Sulawesi Prov., 400 m / 20 km NE Sabbang / 2.28.56S 120.12.00E / 5-7. Jul.2001, Bolm lgt.”. 1 ♂, 1 ♀ in NMPC: “ East Asia Indonesia / S Sulawesi is. / 20 km NE Sabbang / VII-2001 lgt. Bolm ”. 3 ♂♂, 2 ♀♀ in NMPC: “ Indonesia Central Sulawesi / Bonebone 110m / 25-26.V.1996 ” // “ex coll. VÍt Kabourek / National Museum / Prague, Czech Republic” // “ Lophyridia decem- / guttata ( Fabricius, 1801) / det JiřÍ Moravec 2004”. 5 ♂♂, 12 ♀♀ in JWCM: “ CSulawesi / Lore-Lindu NP / Kamorora, 15.4.1994 / leg. R. Gerstmeier et al.”. 4 ♂♂ in JWCM, 1 ♂ in CCJM, 1 ♂ in CJVB: “ C-Sulawesi, Poso-See / 21.- 26.4.1994 / Ponusu b. Tentena / leg. R. Gerstmeier et al.”. 1 ♀ in JWCM: “ 4.4.1995, Indonesia, Tidore, Gurabunga , leg. R. Gerstmeier ”. 1 ♂, 1 ♀♀ in JWCM: “ Indonesia, 15.4.1995 / CSulawesi, Boë / Südende Poso-See / leg. R. Gerstmeier ”. 1 ♂ in JWCM: “4.- 9.7.1999, C Sulawesi, 17 km E Pendolo , 800 m, 120.45.49 E, 2.06.33 S, Bolm lgt.”. 1 ♂, 1 ♀ in JWCM: “5.- 7.7.2001, S. Sulawesi Prov., 400 m, 20 km NE Sabbang , 2.28.56S 120.12.00E, Bolm lgt.”. 1 ♂ in JWCM: “ 17.2.2009, Indonesia, N-Sulawesi / 4 km SE [Pantai] Batu Putih , 150 m / N 1°32’65’’, E 125°07’94[49’] leg. A. Skale ”. 1 ♀ in JWCM: “ 19.2.2009, Indonesia, NSulawesi, 2 km N Airmadidi, S slope Mt. Klabat , 380 m, N 1°24´93´´ E 125°0´66´´, leg. R. Gerstmeier ”. 1 ♂ in JWCM “ 19.2.2009, Indonesia, N-Sulawesi, 10 km N Manado, vic. Wori , 120 m / N 1°34´52´´, E 124°51´60´´ leg. R. Gerstmeier ”. 4 ♂♂, 1 ♀ in JWCM: “ Indonesia, N-Sulawesi / 7 km S Lolak, vic. Bolili / village, 180 m, 22.2. / 2009, leg. R. Gerstmeier / 0°48´65´´, E 124°01´23´´”. 1 ♀ in CDCL: Sulawesi / Palolo, Palu , VII-1986 / Hamdjah & Nishikawa // “Collection / Charles / Dheurle”. 1 ♀ in CDCL: “ Sulawesi. VII-95 / Palolo Palu / Dheurle coll”. 1 ♂ in CDCL: “C. Célèbes. IX.1995 / Palolo Palu / Coll Dheurle ” [printed]. 4 ♂♂ in MHCP, 1 ♂ in MKPC: “c. Sulawesi-Indonesia / 22-28.4 ’98 Rantepao / ( Palopo ) M. Hrdý ”. 1 ♀ in CDCL: “ Sulawesi / Medrano [uncertain locality] I-80. / Schmitt / Collection / Charles / Dheurle”. 2 ♂♂ in CKGC: “ Palopo city environment / Luwu Reg. - Sulawesi Selatan / Indonesia / local collector 06/2017”. 1 ♂, 1 ♀ in CKGC: “ Palolo Palu Area- 1000m / Sulawesi Island / Indonesia / local collector 06/2017”. 1 ♂ in CKGC: “ Palopo city env. 1000m / Sulawesi Selatan Prov. / Indonesia / local collector 06/2017”. 1 ♀ in CKGC: “ Palolo Palu / C – Sulawesi Island / Indonesia / leg. Detani 02/1989”. 1 ♀ in CKGC: “ Palolo environment / Palu Distr. – Tengah prov. / C Sulawesi – Indonesia / local collector 06/2017”. 1 ♀ in CJVB: “ Indonesia IV.1899 / Sulawesi Tengah / Palau Peleng ”. 2 ♂♂ in CKGC: “near Toraut / NSulawesi / Indonesia / local collector 10/2011”. 2 ♀♀ in CKGC: “ 30km NW of Palopo – 700m / Puncak – Sulawesi Island / Indonesia / local collector 06/2010”. 1 ♀ in CKGC: “ Palolo env. - Palu Distr. / Tengah Province / C Sulawesi – Indonesia / local collector 05/2017”. 3 ♂♂ in CKGC: “ Bogani Nani Wartabone NP / Minahassa Peninsula / Sulawesi – Indonesia / leg A. Fritz 10/2012”. 1 ♀ in PSCH: “ Indonesia Sulawesi Barat / 1.4.2012 7 km E Polewali / P. Schüle & N. Rusdi leg. / 03°25.163S 119°24.362E -10”. GoogleMaps 1 ♂ in PSCH: “ Indonesia Sulawesi Barat / 1.4.2012 E Polewali Bakaru / Dam road P. Schüle & N. Rusdi leg. / 03°26.9S 119°32.3E -10a”. GoogleMaps 2 ♂♂, 1 ♀ in PSCH: “ Indonesia Sulawesi Selatan / 8.4.2012 10km N Enrekang / P. Schüle & N. Rusdi leg. / 03°30.708S 119°46.654E -20a”. GoogleMaps 3 ♀♀ in PSCH: “ Indonesia Sulawesi Barat / 6.4.2012 7km E Polewali / P. Schüle & N. Rusdi leg. / 03°25.163S 119°24.362E-17 ”. GoogleMaps 1 ♀ in PSCH: “ Indonesia Sulawesi Selatan / 31.3.2012 9km W Amparita / P. Schüle & N. Rusdi leg. / 04°09.174S 119°47.310E -8”. GoogleMaps 4 ♂♂, 2 ♀♀ in PSCH: “ Indonesia Sulawesi Selatan / 27.3.2012 E Mankara, Berang river / P. Schüle & N. Rusdi leg. / 05°15.878S 119°40.24E -1”. GoogleMaps 1 ♂, 2 ♀♀ in PSCH: “ Indonesia Sulawesi Selatan / 31.3.2012 11km S Batu Batu , / P. Schüle & N. Rusdi leg. / 04°14.768S 119°52.562E -8a”. GoogleMaps 1 ♂ 4 ♀♀ in PSCH: “ Indonesia Sulawesi Selatan / 30.3.2012 ca. 18km E Tanete / P. Schüle & N. Rusdi leg. / 04°31.367S 119°42.320E -6”. GoogleMaps 2 ♀♀ in PSCH: “ Indonesia Sulawesi Selatan / 9.4.2012 S Watanlamuru / P. Schüle & N. Rusdi leg. / 4°33.678S 119°57.533E -26”. GoogleMaps 1 ♂, 1 ♀ in PSCH: “ Indonesia Sulawesi Selatan / 1.4.2012 32.5km SE Polewali / P. Schüle & N. Rusdi leg. / 03°35.909S 119°34.989E -9a”. GoogleMaps 4 ♂♂, 3 ♀♀ in PSCH: “ Indonesia Sulawesi Barat 3.4.2012 / Mamasa road , 13km SW Mamasa / Malabo village P. Schüle & N. Rusdi / leg. 3°03.861S 119°19.278E -14”. GoogleMaps 1 ♂, 1 ♀ in PSCH: “ Palopo, Sulawesi / Indonesia / 05.1992”. 1 ♀ in PSCH: “ Sulawesi, Tengali / S of Palu, Palolo valley / local catcher Jan 1998 ”. 3 ♂♂ in MHCW: “ Pakani, / 38 Km from Palu , / Celebes / 1984, May 12 / Minoru Tao leg.” // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 3 ♂♂ in MHCW: “Kanpong Tuwa [Kampong Tua], / near Palu , / Celebes / 1984, May 13 / Minoru Tao leg. // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 1 ♂, 1 ♀ in MHCW: “ Tokorondo, /near Poso , / Celebes / 1984, May 20 / Minoru Tao leg. // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 1 ♀ in MHCW: “ Ronononcu, / near Poso , Celebes / 1984, May 21 / Minoru Tao leg. // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 2 ♂♂, 3♀♀ in MHCW: “ Kolako , SE Sulawesi, / Indonesia / 2007 / Rudi Hartono leg.”. 5 ♂♂, 1 ♀ in MHCW: “ Lauwo , (at light) / C Sulawesi, / Aug. 7, 1993 / Y. Yamaguchi leg.”. 2 ♂♂, 1 ♀ in MHCW: “ Abuki, / SE Sulawesi, / Indonesia / Apr. 2009 / Rudi Hartono leg.”. 4 ♂♂, 2 ♀♀ in MHCW: Puncak, Palopo , / S Sulawesi, / Indonesia / Nov. 20-23, 2023 / Rudi Hartono leg.”. 1 ♂ in MKPC: “ March 2012 / Indonesia / C. Sulawesi / Puncak Palopo / local collector” // “ Calomera / decemguttata / ( Fabricius, 1801) / Miroslav KlÍcha det.”. 1 ♂ in MKPC: “ X.2011 Indonesia / C. Sulawesi / Toraut / local collector” // “ Calomera / decemguttata / ( Fabricius, 1801) / Miroslav KlÍcha det.”. 5 ♂♂, 4 ♀♀ in MHCW: “Pujalayya [Pajalayya village, east of Benteng ], Selayar Is. , / S Sulawesi, / Indonesia / Feb. 11-15, 2024 / Rudi Hartono leg.”. 1 ♂, 1 ♀ in MHCW: “Batuawu [Watuawu], / near Poso , / Celebes / 1984, May 22 / Minoru Tao leg.” // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 1 ♂, 2 ♀♀ in MHCW: “Dowgara [Dongala], Palu , / Celebes / 1984, May 9 / Minoru Tao leg.” // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 1 ♂, 3 ♀♀ (+ 2 ♀♀ in 100% ethanol) in MHCW: “ Dolo Timur [$?], Palolo, / Palu , C Sulawesi, / Indonesia / Jan. 15, 2024 / Michio Hori leg.”. 1 ♂ in IRSNB: “ Menado [= Manado ] / Coll. Bruin 1877”. 1 ♂, 1 ♀ in IRSNB: “Coll. R. I. Sc. N. B. / Sulawesi Utara / Dumoga-Bone Nat. Park / Picnic Site / 12-X-1985, Station: O23” // “Project Wallace / leg.: R. Bossmans / & J. Van Stalle / I. G. no: 26.977”. 47 paratypes (♂♂, ♀♀) in BMNH: “ Indonesia: / Sulawesi Utara / Dumoga-Bone N. P. ” // “Site 8, 540m / Tumpah Transect / J.D. Holloway 12-13.ii.1985 ” // “R. Ent. Soc. Lond. / Project Wallace / B. M. 1985–10”, some of them irregularly dated as: “January, February, March, August, July, October, November, 1985” and irregularly with additional labels: “Lowland / Rein / forest”, “Lowland forest / edge / ca 200m ”, “ Malaise / trap ”, “ Yellow pan / trap ”, “Banks of R. Tumpah”, “Base camp area / ca 190m ”, or also with: “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 9 ♂♂, 16 ♀♀ in BMNH: “ Lowland / Rein / forest” // “ Sulawesi Tengah: Nr. Morowali, /Ranu River Area , / 27.i.-20.i. v. 1980 ” // “ M.J.D. Brendell / B.M. 1980–280”. 1 ♂ in BMNH: “ Sulawesi Tengah / 1°40´S- 121°20´E / 3-13.iv.1980 ” // “At / light” // “ M.J.D. Brendell / B.M. 1980–280”. GoogleMaps 1 ♀ in BMNH: “ Sulawesi Tengah / 1° 45´S- 121°40´E / 27-30.iii.1980 ” // “At / light” // “ M.J.D. Brendell / B.M. 1980–280”. GoogleMaps 1 ♂ in BMNH: “At / light” // “ Indonesia: / Sulawesi Utara / Manado / 20-21.Nov.85” [handwritten] // “ R. Ent. Soc. Lond. / Project Wallace / B. M. 1985–10” // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 1 ♀ in BMNH: “Site 28, 80m / Coconut-Kayuwatu / J.D. Holloway 10.ix.1985 ” // “ Indonesia: / Sulawesi Utara ” // “ Lophyridia / d. decemguttata / ( Fabricius, 1801) / det. F. Cassola, 1988”. 1 ♂ in BMNH: “ Sangir ” // “10-guttata F. / typical, teste / W. Horn” [handwritten by F. Bates] // “ F. Bates Coll. / 1911–248”. 1 ♂ in BMNH: “ 0°34´36´´N; 123°54´41´´E / alt 215m. / at beach on / R. Tumpah / Sulawesi Utara, Indonesia / 19.X.1985 ”. // W.R.B. Hynd / collection / BMNH(E) 1998–129 ”. GoogleMaps 1 ♂ in BMNH: “ 0°34´36´´N; 123°54´41´´E / alt 215m. / at beach on / R. Tumpah to / 0°34´5´´N;123°54´22´´E / Sulawesi Utara, Indonesia / 28.X.1985 ” // W.R.B. Hynd / collection / BMNH (E) 1998–129”. GoogleMaps 1 ♂ in BMNH: “ 0°34´7´´N; 123°54´32´´E / alt 212m / R. Torauk / Dam – Pagoda / Sulawesi Utara, Indonesia / 8.X.1985 ” // W.R.B. Hynd / collection / BMNH (E) 1998–129”. GoogleMaps 1 ♀ in BMNH: “ 0°34´6´´N; 123°54´21´´E / alt 213 m. / to UV light on / R. Tumpah mouth / 28.X.1985 ”. GoogleMaps 2 ♀♀ in BMNH: “ Celebes / Soputan Masif / Nr. Kelelond. / 14-19.vi.1954 / A.H.G.Alston / B.B. 1954–414” [with reduced elytral maculation]. GoogleMaps 1 ♀ in SDEI: “S. Celebes / Patutuang / Jan. 1896 / H. Fruhstorfer ”. GoogleMaps 1 ♀ in SDEI: “ Samanga / S. Celebes / Nov. 1895 / H. Fruhstorfer ”. GoogleMaps 1 ♂ in SDEI: “ Ost Celebes / Tomboegoe / C. Ribbe, 1882”. GoogleMaps 1 ♀ in SDEI: “ Menado / Nord Celebes / 2de Sem. 1897 / ( Collector C) // “ Coll V. de Poll ”. All paratypes labelled: “ Calomera paradecemguttata sp. nov. / (= C. decemguttata sensu auct.) / det. J. Moravec, C. Dheurle / P. Schüle & J. Wiesner 2024” [red, printed]. Note. Unless otherwise specified, the locality and det. labels are printed. GoogleMaps

Other material examined. 1 ♂ in MKPC: “ Indonesia IV.2000 / Sulawesi ” . 1 ♂ in SDEI: “ Sula Mangoli / Oct.- Novbr. Doherty” . 2 ♀♀ in SDEI: “ Taroena / Groot Sangir 2000 / Doherty III.IV.” . 1 ♀ in SDEI: “ Sangir Is. / Coll. Y. / W. Doherty ” . 1 ♂ in SDEI: “ Laboean / Batjan [= Batan Island ] / Doherty II.III.” // “ Coll V. de Poll ” . 1 ♂ in BMNH: “ Batjan / Aug.-Septbr. / ex Coll. H. Fruhstorfer ” . 1 ♂, 2 ♀♀ in SDEI: “Obi / Doherty” . 1 ♀ in NHMW: “ Lugad / Rosenberg / Poë / Celebes ” . 1 ♂ in IRSNB: “ I.R.Sc. N.B / Obi Island / Indonesia / Sud de Halmahera Island ” // “ Calomera / decemguttata ( Fabricius, 1801) / det F. Cassola 2002” . 2 ♂♂ in PSCH: “ Indonesia, Moluccas centr. / Obi Island – south coast / Tapaya vill. env. 0-300m alt / 20.6.- 20.7.2008 St. Jakl lgt.” . 4 ♂♂ in PSCH: “ Indonesia, C. Moluccas / Obi Isl. – south coast / Seribu Mts. 1200 – 1500m / 22km N of Tapaya vill. / 20.11.- 10.12.2008 St. Jakl leg.” . 2 ♂♂, 1 ♀ in JWCM: “ 14.3.1997, Indonesia / Sulawesi utara ” // “ Pulau Karakelong / Strasse Beo - Rainis / 4 km E Beo, 100m, leg. S. Naumann ” . 2 ♂♂, 1 ♀ in JWCM: “ Indonesia / Sulawesi utara / leg. S. Naumann ” // “Pulau Karakelong / ca 5 km NNW Beo / Einschlag Primärwald / 120 m, 15.3.1997 ” . 2 ♀♀ in JWCM: “ Indonesia / Maluku utara / leg. S. Naumann ” // “Halmahera ( NW) / Baru-Basale / (Stichstrasse), 500m / 5.- 7.3.1997 ” . 2 ♂♂, 2 ♀♀ in JWCM: “ 22.3.1995, Indonesia, Halmahera, 27 km NNW Kao, Tolabit , leg. R. Gerstmeier ” . 3 ♂♂, 3 ♀♀ in JWCM: “ 25.3.1995, Indonesia, Halmahera, Tobelo , leg. R. Gerstmeier ” . 1 ♀ in JWCM: “ 28.3.1995, Indonesia, Halmahera, ca. 15 km W Paca, Wangongira , leg. R. Gerstmeier ” . 2 ♂♂, 4 ♀♀ in JWCM: “ 31.3.1995, Indonesia, Halmahera, ca. 10 km NW Tobelo, Mamuya , leg. R. Gerstmeier ” . 1 ♂ in MFNB: “ Obi Ins. / 1002 / Waterstrad” . 1 ♂, 1♀ in CDCL: “ Ile Gebe / Indonésie / 9. 95 / D. Schmitt ” [with reduced elytral maculation, Fig. 74 View FIGURES 65–74 ] . 1 ♂ in NMPC: “Talaud / Nov. 2011 ” . 4 ♂♂, 6 ♀♀ in NMPC: “ Celebes ” . 1 ♂ in NMPC: “ Indonesia N. Moluccas / Bacan Isl. / 2.12- 5.2.2008, 500–700 m / lgt. Jakl St. ” . 2 ♀♀ in NHMW: “ Java, 1887 / Jakarta [sic!], 4.-15.8. / leg. W. Surkusanang ” // “ Calomera cf / paradecemguttata / Moravec, Dheurle, Schüle / & Wiesner, 2024 in press / det. JiřÍ Moravec 2024 / Locality mislabelled” .

Differential diagnosis. Calomera paradecemguttata sp. nov., hitherto commonly referred to in literature as Calomera decemguttata (sensu auctorum), is externally immediately distinguished from the true Cicindela decemguttata by the pattern of its yellowish elytral maculation ( Figs48,50–51 View FIGURES 48–51 , 65–74 View FIGURES 65–74 ), particularly with lateromedian macula adjacent to the outer elytral margin and in form of continuous, cranked lateromedian-discal band (in contrast to the lateromedian macula in C. decemguttata , which is distinctly or always at least noticeably separated from the outer elytral margin ( Figs 1–2 View FIGURES 1–3 , 4–5 View FIGURES 4–5 , 11–19 View FIGURES 11–19 ). Other diagnostic characters include at least partly densely setose genae and proepisterna ( Fig. 55 View FIGURES 52–55 ), as well as notably coarser pronotal surface sculpture with usually numerous setae along dorsolateral discal areas ( Figs 52–54 View FIGURES 52–55 ), in contrast to the only partly and sparsely setose proepisterna, almost glabrous genae (usually with only few setae at their base), finer pronotal surface sculpture with sparser discal dorsolateral setosity in C. decemguttata ( Figs 6–9 View FIGURES 6–10 ), even if the setae on the surfaces are easily abraded and the setosity sometimes varies. Discal-juxtasutural catoptric area on female elytra ( Figs 51 View FIGURES 48–51 , 68–69 View FIGURES 65–74 ) is mostly distinct. Males of C. paradecemguttata sp. nov. are clearly distinguished by the shape of the aedeagus apical portion in its dorsal (and ventral) aspect having on each side of its lateral edges characteristic indentation ( Figs 76, 78, 80, 82, 84, 88 View FIGURES 75–92 ), in contrast to the aedeagi of C. decemguttata (and all others of this species-complex) which have their lateral edges in dorsal (and ventral) view continuously conically attenuated towards apex ( Figs 32, 40 View FIGURES 31–41 ).

Description. Body ( Figs 48, 50–51 View FIGURES 48–51 ) variably medium-sized to large, 11.1–13.5 (HT 12.4, AT 11.8) mm long, 4.20–5.05 (AT 4.50) mm wide (females usually larger but sometimes smaller than males, exceptionally (from Obi Island) 14.2 mm long, 5.25 mm wide.

Head ( Figs 48, 50, 51 View FIGURES 48–51 , 56–58 View FIGURES 56–64 ), with large eyes but notably narrower than body (across elytra), width 3.10–3.60 (–3.80) mm; all dorsal portions glabrous.

Frons fluently passing into vertex, its surface rather distinctly convex in middle, black-coppery or more vividly cupreous, extremely finely and densely longitudinally striate, striae becoming irregularly asperate in middle when passing on vertex; supraantennal plates rather large, flat, metallic black-green, or variably iridescent green, green-blue, reddish-cupreous or violet.

Vertex almost flat with small, limited convection in middle, dark coppery or vividly iridescent cupreous, often (mostly in males) with limited blue-green or violet-blue anterior-sublateral stripes, in females more often uniformly coppery yet sometimes with vividly cupreous, rarely greenish iridescence, very rarely almost black; anteromedian area extremely finely irregularly granulate-asperate to extremely irregularly finely rugulose (as passing from frons) becoming very irregularly rugulose and wavy posteriad; juxtaorbital areas densely parallel striate yet striae on juxtaorbital margin sometimes irregularly fragmented, while more regular and thicker on sublateral areas when divergent posteriad as passing onto temples and genae; postero-median area irregularly vermicular-rugulose, sculpture passing onto notably convex occiput (mostly hidden under pronotum).

Genae ( Fig. 55 View FIGURES 52–55 ) iridescent green, sometimes with bronze-cupreous lustre, finely and densely parallel striate, primarily rather densely whitish setose, either prevailingly on their posteromedian area, or at least on posteroventral area, yet the setae are easily (usually partly) abraded; anterior area almost glabrous.

Clypeus variably deep metallic green, green-blue or cupreous with green-blue margins, alternatively darkviolet, surface glabrous, finely irregularly wrinkled or coriaceous.

Labrum ( Figs 59–64 View FIGURES 56–64 ) primarily 6-setose (occasionally with five or seven setae developed); male labrum almost uniformly ivory-yellow or yellow to ochre-testaceous with only narrow blackish margins and blackened teeth, mostly almost uniformly shaped in both sexes and comparatively rather long, length 0.85–1.05 (–1.15) mm, width 1.70–1.85 (–2.00) mm; central convexity moderate; basolateral margins only slightly convex, basolateral anterolateral teeth absent or bluntly indicated; anterior margin almost uniformly tridentate, the teeth right-angled but primarily pointed, median tooth variably slightly or notably longer.

Mandibles ( Figs 48, 50–51 View FIGURES 48–51 , 56–58 View FIGURES 56–64 ) sexually dimorphic in coloration: ivory-yellow or ochre-testaceous with black teeth in male, while metallic black with only limited testaceous basal area in female, normally shaped with arcuate lateral margins, almost symmetrical, both mandibles with four teeth (and basal molar), apical teeth of moderate length (comparatively short within the genus); second tooth larger than third and fourth teeth, yet slightly variably the third tooth in left mandible is sometimes smaller than the fourth.

Palpi ( Figs 56–58 View FIGURES 56–64 ). Maxillary palpi with longest palpomere in males variably metallic black-brown, or brownish-testaceous (also in HT Fig. 56 View FIGURES 56–64 ), while constantly black-brown or metallic black in females, penultimate and terminal palpomeres metallic black in both sexes, in males mostly with green, green-blue or reddish-cupreous lustre; labial palpi in both sexes with longest palpomere yellow-testaceous (often darkened in female), terminal palpomere metallic black.

Antennae ( Figs 48, 50–51 View FIGURES 48–51 , 56–58 View FIGURES 56–64 ) rather long, almost reaching elytral half in males, elytral third in females; antennomeres 1–4 metallic-black, usually with strong greenish-blue, rarely partly faint reddish-cupreous lustre, in females generally darker (in some males from Obi Island entirely iridescent reddish-cupreous), with sparse stiff setae; antennomeres 5–11 greyish-blackened with usual micropubescence.

Thorax. Pronotum ( Figs 48, 50–55 View FIGURES 48–51 View FIGURES 52–55 ), slightly shorter than wide, length 2.40–2.50 (–2.80) mm, width 2.50– 2.60 (–3.00) mm; rather variably coloured, mostly dark coppery with diffusing reddish-coppery lustre and stronger reddish lustre on anterior and posterior lobe and limited green-blue or violet lustre within sulci, or the greenish coloration and iridescence prevailing (also in HT, Fig. 52 View FIGURES 52–55 ); anterior lobe of almost the same width as posterior lobe (but both are narrower than disc), its surface very irregular and rugulose of an indefinable pattern; anterior sulcus well pronounced; disc with lateral margins moderately convex in both sexes yet in females often subparallel or moderately or more markedly attenuated posteriad; notopleural sutures visible from above as parallel with pronotal lateral edges and proepisternal outer margins; discal surface rather coarsely, very irregularly rugulose of barely definable pattern except for deeper, transverse-parallel rugae along mostly distinctly pronounced median line; posterior lobe almost smooth, or irregularly with indistinct and indefinite sculpture, its dorsolateral bulges rather distinctly pulvinate; posterior rim indefinite, sometimes consisting of two or three transverse wrinkles; proepisterna ( Fig. 55 View FIGURES 52–55 ) metallic coppery with various, mostly green-blue lustre, their surface finely coriaceous-asperate and with few parallel juxtanotopleural wrinkles, primarily rather densely covered with appressed, rather long whitish setae, covering at least two thirds of proepisternal surface; mesepisterna ( Fig. 55 View FIGURES 52–55 ) lustrously metallic black-green or shiny metallic green, glabrous, except for a cluster of white setae at ventral rim; female mesepisternal coupling sulci distinct, in form of deep shortly longitudinal impression; metepisterna (partly obvious in Fig. 55 View FIGURES 52–55 ) metallic black-blue or metallic green, sometimes with reddish-cupreous lustre, surface coriaceous, densely appressed setose; prosternum and mesosternum mostly iridescent green-blue almost smooth or coriaceous, glabrous except for several lateral setae; metasternum metallic black with strong green-blue lustre, smooth, glabrous in middle, densely whitish setose on lateral areas.

Elytra ( Figs 48, 50–51 View FIGURES 48–51 , 65–74 View FIGURES 65–74 ) elongate, 7.10–8.55 (–9.15) mm long, dorsally moderately convex, humeral impressions short and shallow yet recognizable, often due to an inside-arranged clump of faintly iridescent setigerous punctures, basodiscal convexity moderate, discal impression indistinct, practically absent; apical impressions moderate; elytral shape slightly sexually dimorphic: humeri rounded in both sexes, outer lateral margins in males moderately convex, in females usually notably more dilated in middle; anteapical angles arcuate towards subacute apices in males, while apices rounded in females; sutural spine short and indistinct; microserrulation irregular, very fine yet clearly obvious; epipleura normally shaped; elytral punctation throughout, yet indistinct due to extremely shallow and densely arranged, dark green punctures; anterior elytral half may appear micro-granulate and limited antero-discal area indefinitely yet more coarsely irregularly tuberculate; basodiscal area possessing sparse, irregular and sometimes inconspicuous rows of reddish and green, faintly iridescent setigerous foveae, yet whitish setae arising from the foveae sparse and easily abraded; other elytral parts almost glabrous except for few juxtaepipleural setae; sutures often in form of more or less distinct faintly iridescent cupreous or blue-green, sometimes darkened stripes (depending upon illumination angle), yet also distinctly iridescent green (as in HT, Fig. 48 View FIGURES 48–51 ); discal-juxtasutural catoptric area on female elytra ( Figs 51 View FIGURES 48–51 , 68–69 View FIGURES 65–74 ) mostly distinct, conspicuously iridescent green or reddish-cupreous, consisting of densely arranged light-reflecting punctures; elytral coloration notably matt, appearing velvety black, particularly on elytral disc, rarely with only indistinct cupreous iridescence, more often with olivaceous-green lustre diffusing particularly along anteapical angles; surface glabrous (except for the few and mostly indistinct or abraded setae arising from the foveae on basodiscal convexity) and sparsely present darker setae along epipleura; elytral maculation laterally adjacent to outer elytral margin, whitish or ivory-yellow, formed primarily by six maculae, yet the lateromedian macula is always connected with central macula in form of mostly continuous, cranked lateromedian-discal band; humeral macula separated from variably shaped subhumeral macula; small, elongate lateral spot placed underneath the lateromedian-discal band is sometimes connected with the band by a thin, juxtaepipleural stripe; anteapical-apical lunule always present, with anteapical mesad-directed protrusion and running along the anteapical-apical margin but mostly separated from apical suture; anomalous adults with extremely reduced maculae ( Fig. 74 View FIGURES 65–74 ) are obviously very rare (see “Variability” and “Distribution” below).

Abdomen. Ventrites metallic black usually with strong green-blue lustre (depending upon illumination angle), densely covered with whitish, rather long and appressed setae, which are densest on first three or four visible ventrites, becoming much sparser posteriad, last two ventrites sometimes almost glabrous.

Legs. Coxae metallic black with green-blue lustre, pro- and meso-coxae with cluster of whitish setae at base; metacoxae glabrous with only sensory seta in middle, while lateral areas with cluster of whitish (easily abraded) setae; trochanters in both sexes shiny black-brown, metatrochanters almost black, first of the pro- and mesotrochanters with indistinct subapical seta; femora dorsally metallic black with only faint green-blue lustre, in ventral view usually (also in HT) with strong shiny green-blue lustre; profemora densely covered with rows of whitish, mostly erect setae, which are somewhat sparser on mesofemora except for long and dense setae on their basoventral area, sparser and shorter but stiffer on metafemora; tibiae with two (easily broken) thorn-like apical setae, dorsally metallic-black, protibiae often with greenish or reddish-cupreous lustre, which is obvious on ventral area of meso- and metatibiae (also in HT); pro- and mesotibiae covered with rows of scattered, short whitish setae, mesotibiae also with usual, dense pad of whitish and greyish setae on their apical half; metatibiae with sparse, short, stiff to almost thorn-like rusty setae; tarsi metallic black, usually with more or less strong green-blue lustre; as usual, the first three protarsi in males distinctly dilated; claws black-brown.

Aedeagus ( Figs 75–92 View FIGURES 75–92 ) elongate, 3.50–3.85 mm long, 0.80–0.85 (–0.95) mm wide, widest in middle and then in subapical ventral outline, then conically constricted towards apex which is in its left lateral view of a rather variable shape (as in most others of the species-complex), appearing bilobed as consisting of a narrowly conical, more sclerotized blunt knob and a dorsal, rounded nose-like lobe; in dorsal (and ventral) view ( Figs 76, 78, 80, 82, 84, 88 View FIGURES 75–92 ) the lateral edges of the apical portion possessing distinct indentations (on each side of the apical portion). Internal sac ( Figs 85–86, 89–92 View FIGURES 75–92 ) widely armed, with conspicuous arciform piece which in right lateral view ( Figs 86, 90 View FIGURES 75–92 ) appears as conspicuous central spike; as in other species the internal sac possesses a multi-convoluted flagellum associated with sustaining membranous “auricular” sheets; the loops of the convoluted flagellum change their shapes when the aedeagus is somewhat turned and appears differently shaped in the right lateral view. Nevertheless, as mentioned in “Variability”, “Remarks” and “Distribution” below, the internal sac in the male from Obi Island, Maluku, appears different.

Variability. All type specimens of this new species possess usual variability in coloration of the head and pronotum and insignificant variability in whitish elytral maculation, as treated in the description above, except for the aberrant adults with reduced maculation, such as from Palau Peleng ( Fig. 73 View FIGURES 65–74 ). One historical male specimen with notably reduced elytral maculation (paratype in SDEI), labelled “Tombaogoe” (= Tombugu, also spelled Tombuku), comes from the Central Sulawesi Province, Marowali Regency. Some specimens from other islands of the Maluku, such as the Gebe Island (CDCL, Fig. 74 View FIGURES 65–74 ) and two females (SDEI) from Taroena, Mt. Sangir in the Sangihe Islands (also spelled Sangir) north-east of the Sulawesi Minahassa Peninsula, also possess considerably reduced maculation. The aedeagi of males with such reduced elytral maculation possess the same shape as in the holotype and all other examined males of C. paradecemguttata sp. nov., including their lateral indentations in dorsal view. However, males from Obi Island, such as the male illustrated by Schüle (2010, fig. 5) under the at-the-time-confused-name C. decemguttata , although possessing aedeagi as in C. paradecemguttata sp. nov., their internal sac ( Fig. 92 View FIGURES 75–92 ) somewhat differs. Added to that, as also obvious from the above-mentioned fig. 5 by Schüle (2010), the pronotal surface has finer sculpture and is more reddish, antennomeres 1–4 iridescent red, and elytral maculation wider in most of the specimens from Obi Island than in the type specimens; moreover, several females from Obi Island are notably larger and with wider elytral maculation, thus resembling C. mamasa (their measures are in parenthesis in the “Description” above). See also “Remarks” and “Distribution” below. In some very old male specimens, the dorsal aspect of the aedeagus has the lateral indentation obvious on only one side, as it was probably partly deformed when extracted from the abdomen of the hard body supposedly killed by strong ethanol.

Distribution. Calomera paradecemguttata sp. nov. inhabits the large Indonesian island Sulawesi and some other islands of Maluku. The type locality is Wakarumba on the island of Palau Buton adjacent to the southeastern peninsula of Sulawesi. All other type specimens also come from the main Sulawesi territory which is formed also with small adjacent islands. Most adults were taken in the area of Palu Palolo Valley southeast of Palu, mostly by local insect dealers. Palu, despite its northwestern location, is the capital of the (administrate) Central Sulawesi Province (Sulawesi Tengah). In the Polewali Mamasa Regency , Province of West Sulawesi (Sulawesi Barat), the new species is sympatric, but not syntopic with C. mamasa as inhabiting different areas of lower altitudes. The third author of the present paper collected C. paradecemguttata on riverbanks of the Jeneberang river (“Berang” on the labels) in South Sulawesi, east of Mankara ( Fig. 236 View FIGURE 236 ), as well as near the Mamasa river. This species has obviously a rather wide range of habitats, such as sea beaches, but also away from the water, occurring also on roads and other biotopes, and almost year-round (depending on altitudes). Adults from Rantepao, Palopo (MHCP) were taken on a stony ground of a quarry near a road through a hilly area, outside a forest. Some paratypes come from northern Sulawesi areas, Manado (also spelled “Menado”) and from the Minahassa Peninsula with the Bogani Nani Wartabone (= Dumoga Bone) National Park. The specimens occurring in North Maluku, Halmahera, on the island of Obi, island of Bacan and other islands, including the Sangihe Islands north-east of the Minahassa Peninsula, as well as far more distant Palau Karakelang, the main island of the Talaud Islands north east of Sulawesi, are not included as paratypes, particularly due to the great biodiversity in Indonesia and possible allopatric speciation (see also “Variability” above).

The two above-listed female specimens with“ Java,Jakarta ”on their labels have characters of C. paradecemguttata sp. nov., but were evidently mislabelled; an accidental spreading from Sulawesi, e.g. by ships, is conceivable yet in this case of the only two specimens appears improbable.

It must be emphasized here that in accordance with the above-discussed confusion, specimens listed by Cassola (1991) from Sulawesi under C. decemguttata (including those listed here from MCHW) were in fact C. paradecemguttata sp. nov.

Remarks. Despite the rather homogenous characters (except for exceptionally aberrant adults), particularly the shape of the aedeagus apex in its dorsal (and ventral) aspect, which immediately differentiate C. paradecemguttata sp. nov. from C. decemguttata and others of the species complex, we have included into the type series only specimens from the Sulawesi main territory, which includes adjoining islands (for the reasons see “Variability” and “Distribution” above.