Calledema, BUTLER, 1878

6. CALLEDEMA BUTLER, 1878 View in CoL

Calledema Butler, 1878 . Trans, ent. Soc. Lond: 65. Type species: Calledema marmorea Butler, 1878 . ibid.: 65.

Adults. ( Figure 1F View FIGURE 1 , 10D View FIGURE 10 ) Head: male and female antennae simple filiform; antennal tufts present; third segment of labial palpus long; ocelli present. Thorax: male prothoracic leg with scent pocket; tarsal claws bifid. Wings: males 33 to 49 mm, females 37 to 55 mm; forewing pattern with pastel coloration or dark brown with contrasting metallic markings; accessory cell present; hindwing with Sc-Rs stalked. Abdomen: hair-like terminal tufts present, fifth abdominal sternite with peniculus in some species ( Figure 4J View FIGURE 4 ). Male terminalia ( Figure 6A View FIGURE 6 ): eighth tergite with midplate; eighth sternite with r-shaped antecosta, caudal edge with v-shaped indentation, slightly to extremely asymmetrical. SSO well-developed; valvae costa sclerotized but apically membranous (long ear shaped projections arising from the outer side of each valve); uncus dorsally arrowhead-like, ventrally hook-like; massive sclerotized socii; juxta sclerotized and fused to the sacculus, costulae may be present (cattle hoof shape in C. humboldti (Thiaucourt) and C. rufescens (Schaus)) . Aedeagus long and thin, the distal portion somewhat expanded, apex with small thorns/deciduous cornuti. Female terminalia: Papillae anales moderately to extremely sclerotized; dorsal sclerotized scent organ presents between papillae anales. Eighth tergite usually massive, with lateral prominences reduced or absent, anterior apophyses variable. Eighth sternite similar to tergite; ostium bursae wide. Ductus bursae sclerotized and dorso-ventrally flattened, with sclerotized left-hand projection. Corpus bursae with a sclerotized shield, otherwise membranous; signum and sclerotized invaginations sometimes present on shield.

Diagnosis. The species of this genus can be easily recognized due to the presence of inverted triangles on the costa, and a contrasting transverse line or band on the forewing, or discal dashes and subterminal lines giving the overall patterning a complex appearance. The presence of the peniculus on the fifth abdominal sternite, and the combination of the hook-like uncus and massive socii, are also unique to this genus.

Comments. Calledema is a composite Greek word, “Calli-Callo” translates as “beautiful” and “demeo” as “to build”. Thus, Calledema means “beautifully built” ( Weller 1989). Host plants include Chrysobalanaceae ( Microdesmia arborea , Hirtella triandra , Hirtella racemosa , Hirtella guatemalensis , Licania hypoleuca ), Combretaceae ( Combretum farinosum , Terminalia amazonia), Melastomataceae ( Miconia stevensiana ), and Myrtaceae species ( Eugenia basilaris , Myrcia splendens ) (ACG database).

Distribution. From Mexico to Argentina.

Species included (14).

Calledema arema Schaus, 1905

Calledema argenta Schaus, 1905

Calledema contingata (Möschler, 1883)

Calledema didugana (Draudt, 1932)

Calledema electra Thiaucourt, 1985

Calledema humboldti (Thiaucourt, 1985)

Calledema jocasta Schaus, 1901

Calledema marmorea Butler, 1878 *

Calledema plusia (Felder, 1874)

Calledema plusioides (Felder, 1874)

Calledema ronaldi Schaus, 1934

Calledema rufescens (Schaus, 1905)

Calledema sodalis Butler, 1878

Calledema sura Schaus, 1905