Lysana, MOSCHLER, 1883

24. LYSANA MÖSCHLER, 1883 View in CoL

Lysana Moeschler, 1882 . Verhandl. zool.-bot. Ges Wien 32: 347. Type species: Lysana plexa Moeschler, 1882 . ibid. 32: 347.

Adults. ( Figure 3A View FIGURE 3 , 4I View FIGURE 4 , 11C View FIGURE 11 ) Head: male and female antennae simple filiform; antennal tufts present, in some species these are extremely long and extend well above the head; third segment of labial palpus long. Thorax: male prothoracic leg with double scent pocket on femur, a large, membranous, glandular area on tibia, and an elaborate scent pencil; tarsal claws simple; ocelli present. Wings: males 37 mm, females 44 mm; females with dark hindwing; males with white hindwing; forewing with accessory cell present, hindwing with Sc-Rs stalked. Abdomen: male second abdominal sternite with long slender apodemes. Male terminalia ( Figure 6O View FIGURE 6 ): eighth tergite with midplate; eighth sternite plain. SSO present, valvula not pleated, with many long hair-like scales and setae; costa sclerotized entire length. Uncus tip sclerotized; socii sclerotized, thin and setose. Costulae absent; juxta deeply concave. Aedeagus callosum foot-like; long, slender and slightly curved. Female terminalia: Papillae anales membranous with short, setae. Eighth tergite caudal edge rounded with slightly incurved, lateral processes. Eighth sternite caudal edge slightly rounded, with lateral thickenings extending from ostium bursae to caudal edge; ostium bursae straight and unornamented. Ductus bursae proximal third sclerotized, next third tubular with sclerotized patches; distal third membranous. Corpus bursae-membranous, lacking defined signum; small lightly sclerotized patch of spinules extending ventrally.

Diagnosis. The complex pattern and coloration of the type species is unique. The remarkably long scales on the antennae are a prominent feature of members of this genus. The forewings of the type species have a single, white, rounded triangle in the upper corner of the discal cell, other species are more darkly patterned with complex ornamentation. The large papillae anales on the female genitalia are unique, as well as the large and sclerotized then membranous large ductus bursae. The modified second segment, the irregularly folded saccular edge of the valvae and a simple triangular uncus are in combination rather unique.

Comments. In males, the shape of the costal flange differs slightly as does the shape of the costal tip. We do not consider these minor variations in shape and size to constitute species-specific differences. Females of L. plusiana have long villar projections lining the entire inside of the corpus bursae, and this condition is unique to the species ( Weller 1989). Compared to other nystaleines, Lysana plusiana has extremely long antennal tufts. Becker (2014) incorrectly listed the name parvipuncta Dognin, 1909 in two places in his checklist: as a synonym of Poresta lanassa (Druce) and as a valid species of Lysana . The proper assignment is as a synonym of P. lanassa , since parvipuncta does not belong to Lysana . Female genitalia illustrated in Weller (1989).

Distribution. From Costa Rica to Brazil.

Species included (6).

Lysana abnormis (Schaus, 1905)

Lysana argolarma (Draudt, 1932)

Lysana improvisa (Dognin,1923)

Lysana minasensis Schaus, 1928

Lysana plexa Möschler, 1883 *

Lysana plusiana Schaus, 1901