Atheta ( Dimetrota auct .) hampshirensis Bernhauer, 1909

Atheta ( Dimetrota auct.) hampshirensis Bernhauer, 1909 View in CoL

( Figs. 13–14 View FIGURES 10–14 in Lohse & Smetana 1985; Figs. 71–74 View FIGURES 70–74 in Klimaszewski & Winchester 2002)

Homalota moesta Mäklin View in CoL in Mannerheim, 1852: 307 ( nec Zetterstedt, 1828: 101).

Atheta ( Dimetrota) moesta: Bernhauer, 1907: 394 View in CoL (as valid species).

Atheta ( Dimetrota) hampshirensis Bernhauer, 1909: 525 View in CoL .

Atheta ( Philhygra) leviceps Casey, 1910a: 66 , syn. nov.

Dimetrota ( s. str.) sectator Casey, 1910a: 102 View in CoL , syn. nov.

Dimetrota ( s. str.) retrusa Casey, 1910a: 103 View in CoL , syn. nov.

Datomicra ( s. str.) hebescens Casey, 1910a: 124 View in CoL , syn. nov.

Datomicra ( s. str.) insolida Casey, 1910a: 125 View in CoL , syn. nov.

Datomicra ( s. str.) pellax Casey, 1910a: 124 View in CoL , syn. nov.

Pseudota vana Casey, 1911a: 150 View in CoL , syn. nov.

Dimetrota sectator: Casey, 1911b: 252 View in CoL (as valid species).

Dimetrota insolida: Casey, 1911b: 252 (as synonym of Dimetrota sectator View in CoL ).

Atheta ( Dimetrota) maeklini Fenyes, 1920: 205 (replacement name for Atheta moesta View in CoL (Mäklin in Mannerheim, 1852) ( nec Zetterstedt, 1828).

Atheta ( Dimetrota) moesta: Fenyes, 1920: 205 View in CoL (as synonym of At. maeklini ).

Atheta ( Pancota) vana: Fenyes, 1920: 203 View in CoL (as valid species).

Atheta ( Dimetrota) hampshirensis: Fenyes, 1920: 205 View in CoL (as valid species).

Atheta ( Dimetrota) sectator: Fenyes, 1920: 205 View in CoL (as valid species).

Atheta ( Dimetrota) insolida: Fenyes, 1920: 205 View in CoL (as synonym of At. sectator View in CoL ).

Atheta ( Dimetrota) retrusa: Fenyes, 1920: 205 View in CoL (as valid species).

Atheta ( Datomicra) hebescens: Fenyes, 1920: 220 View in CoL (as valid species).

Atheta ( Datomicra) pellax: Fenyes, 1920: 220 View in CoL (as valid species).

Atheta ( Pancota) vana: Bernhauer & Scheerpeltz, 1926: 661 View in CoL (as valid species).

Atheta ( Dimetrota) hampshirensis: Bernhauer & Scheerpeltz, 1926: 663 View in CoL (as valid species).

Atheta ( Dimetrota) maeklini: Bernhauer & Scheerpeltz, 1926: 664 (as valid species).

Atheta ( Dimetrota) moesta: Bernhauer & Scheerpeltz, 1926: 664 View in CoL (as synonym of At. maeklini ).

Atheta ( Dimetrota) sectator: Bernhauer & Scheerpeltz, 1926: 665 View in CoL (as valid species).

Atheta ( Dimetrota) insolida: Bernhauer & Scheerpeltz, 1926: 665 View in CoL (as synonym of At. sectator View in CoL ).

Atheta ( Dimetrota) retrusa: Bernhauer & Scheerpeltz, 1926: 665 View in CoL (as valid species).

Atheta ( Datomicra) hebescens: Bernhauer & Scheerpeltz, 1926: 668 View in CoL (as valid species).

Atheta ( Datomicra) pellax: Bernhauer & Scheerpeltz, 1926: 668 View in CoL (as valid species).

Atheta ( Dimetrota) hampshirensis: Moore & Legner, 1975: 362 View in CoL (as valid species).

Atheta ( Dimetrota) maeklini: Moore & Legner, 1975: 365 (as valid species).

Atheta ( Dimetrota) moesta: Moore & Legner, 1975: 365 View in CoL (as synonym of At. maeklini ).

Atheta ( Dimetrota) sectator: Moore & Legner, 1975: 373 View in CoL (as valid species).

Atheta ( Dimetrota) insolida: Moore & Legner, 1975: 373 View in CoL (as synonym of At. sectator View in CoL ).

Atheta ( Dimetrota) retrusa: Moore & Legner, 1975: 372 View in CoL (as valid species).

Atheta ( Datomicra) hebescens: Moore & Legner, 1975: 362 View in CoL (as valid species; misspelled as herbescens).

Atheta ( Datomicra) pellax: Moore & Legner, 1975: 369 View in CoL (as valid species).

Atheta ( Pancota) vana: Moore & Legner, 1975: 376 View in CoL (as valid species).

Dimetrota hampshirensis: Seevers, 1978: 259 View in CoL (as valid species).

Dimetrota maeklini: Seevers, 1978: 259 (as valid species).

Dimetrota moesta: Seevers, 1978: 259 (as synonym of Dim. maeklini ).

Dimetrota sectator: Seevers, 1978: 260 View in CoL (as valid species).

Dimetrota insolida: Seevers, 1978: 260 (as synonym of Dim. sectator View in CoL ).

Dimetrota retrusa: Seevers, 1978: 260 View in CoL (as valid species).

Datomicra hebescens: Seevers, 1978: 259 View in CoL (as valid species).

Datomicra pellax: Seevers, 1978: 259 View in CoL (as valid species).

Pseudota vana: Seevers, 1978: 261 View in CoL (as valid species).

Dimetrota moesta: Lohse & Smetana, 1985: 289 (as valid species).

Atheta ( Dimetrota) moesta: Klimaszewski & Winchester, 2002: 29 View in CoL (as synonym of At. hampshirensis View in CoL ).

Atheta ( Dimetrota) hampshirensis: Klimaszewski & Winchester, 2002: 29 View in CoL (as valid species).

Atheta ( Dimetrota) maeklini: Klimaszewski & Winchester, 2002: 29 (as synonym of At. hampshirensis View in CoL ).

Type material. Lectotype of Homalota moesta (designated by Lohse and Smetana (1985)): UNITED STATES: Alaska: ♂, Sitka ( MZHF).

Lectotype of Atheta hampshirensis (here designated): ♂, “N.[ew] H.[ampshire], Fall”, “7603”, “ hampshirensis Bernh. Typus. Fenyes!” (yellow label), “Chicago NHMus M.Bernhauer Collection ” ( FMNH).

Lectotype of Atheta leviceps (here designated): ♀, “N.[ew] Y.[ork] [ Catskill Mts. ]”, “ leviceps Csy. ”, “TYPE USNM 39362” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotypes: 2♂♂, ♀, “N.Y.[ Catskill Mts. ]”, “ leviceps PARATYPE USNM 39362” (red label), “CASEY bequest 1925” ( NMNH) .

Lectotype of Dimetrota sectator (here designated): ♂, “Cal.[ifornia] [ Gilroy Hot Springs , Santa Clara Co.]”, “ sectator Csy. ”, “TYPE USNM 39118” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotypes: 2♂♂, 2♀♀, specimens, “Cal.[ifornia] [ Gilroy Hot Springs , Santa Clara Co.]”, “ sectator PARATYPE USNM 39118” (red label), “CASEY bequest 1925” ( NMNH) .

Lectotype of Dimetrota retrusa (here designated): ♂, “Metlakatla, B.[ritish] Col.[umbia]”, “ retrusa Csy. ”, “TYPE USNM 39120” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotypes: ♂, 5♀♀, “Metlakatla, B.[ritish] Col.[umbia]”, “ retrusa PARATYPE USNM 39120” (red label), “CASEY bequest 1925” ( NMNH) .

Lectotype of Datomicra hebescens (here designated): ♂, “N.[ew]Y.[ork][ Catskill Mts. ]”, “ hebescens Csy. ”, “TYPE USNM 39176” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotypes: 8 specimens, “N.[ew]Y.[ork][ Catskill Mts. ]”, “ hebescens PARATYPE USNM 39176” (red label), “CASEY bequest 1925” ( NMNH) .

Lectotype of Datomicra insolida (here designated): ♂, “Cal.[ifornia] [ Gilroy Hot Springs , Santa Clara Co.]”, “ insolida Csy. ”, “TYPE USNM 39119” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotypes: 2 specimens, “Cal.[ifornia] [ Gilroy Hot Springs , Santa Clara Co.]” , 7 specimens (1 without head and prothorax), “Cal.[ifornia] [San Francisco]”, “ insolida PARATYPE USNM 39119” (red label), “CASEY bequest 1925” ( NMNH) .

Holotype of Datomicra pellax : ♀, “R.[hode] I.[sland][Boston Neck]”, “ pellax Csy. ”, “TYPE USNM 39171” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Pseudota vana (here designated): ♂, “Q.[ueen] C.[harlotte] I.[slands][Massett (Keen)]”, “ vana Csy. ”, “TYPE USNM 39157” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotype: ♀, “Q.[ueen] C.[harlotte] I.[slands][Massett (Keen)]”, “ vana PARATYPE USNM 39157” (red label), “CASEY bequest 1925” ( NMNH) .

Additional material: UNITED STATES: Alaska: ♀, Kenai ( Holmberg) ( MZHF) ; New York: Orange Co .: 4 specimens, 8 km S New Windsor, Black Rock Forest , 23– 25.v.1998 ( V.I.Gusarov) ( SPSU) ; North Carolina: Haywood Co .: 13 specimens, Cold Springs Creek, Forest Road 148, 29 km N Waynesville, Pisgah National Forest , 35º44.6'N 83º00.3'W, 750 m, 4.vi.2001 ( V.I.Gusarov) ( SPSU) GoogleMaps ; Pennsylvania: ♀, 41º08.39'N 75º32.24'W, in mushrooms, 16.viii.1998 ( V.I.Gusarov) ( SPSU) GoogleMaps ; CANADA: Québec: ♂, St.­Jacques­de­Leeds , 46º16'N 71º23'W, maple forest, 8.vii.1994 ( SPSU) GoogleMaps .

Diagnosis. See Lohse & Smetana (1985).

Discussion. The types of Homalota moesta , At. hampshirensis , At. leviceps , Dim. sectator , Dim. retrusa , Dat. hebescens , Dat. insolida , Dat. pellax and Pseudota vana are similar in external characters and the shape of the aedeagus and spermatheca.

Distribution. Atheta hampshirensis is widespread in the United States ( Alaska, Washington ( Lohse & Smetana 1985), Oregon, California, New Hampshire, Rhode Island, New York, Pennsylvania and North Carolina) and Canada ( British Columbia and Québec).