Soderstrom, , N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT, N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT

384 L SÖDERSTRÖM, N.A. KONSTANTINOVA, A. HAGBORG, T. PRESTØ, K. HASSEL & M. VON KONRAT

** Jungermannia eucordifolia Schljakov. View in CoL SVALBARD: Düll 1983 as Jungermannia exsertifolia View in CoL . NOTE: Düll’s (1983) report is erroneous (cf. Frisvoll & Elvebakk, 1996).

Lophozia (Dumort.) Dumort. View in CoL

*** Lophozia guttulata (Lindb. et Arnell) A.Evans. View in CoL SVALBARD: Frye & Clark 1945 as Lophozia porphyroleuca, Smith 1990 as Lophozia longiflora . BJØRNØYA: Watson 1922 as Lophozia porphyroleuca, Summerhayes & Elton 1923a as Lophozia porphyroleuca . NOTE: The occurrence of this almost strictly xylicolous species is highly unlikely and Frisvoll & Elvebakk (1996) rejects it. Some reports are probably due to nomenclature confusions (cf. Söderström et al., 2013).

* Lophozia schusteriana Schljakov. View in CoL SVALBARD: Schuster 1969 as Lophozia groenlandica . HAAKON VII LAND: Arnell & Mårtensson 1959 as Lophozia groenlandica . OSCAR II LAND: Arnell & Mårtensson 1959 as Lophozia groenlandica . NOTE: The name Lophozia groenlandica is in a few cases connected with Lophozia schusteriana View in CoL but this should probably not be the case of any European material, which instead should go to Lophozia murmanica View in CoL or Lophozia wenzelii View in CoL .

Lophoziopsis Konstant. et Vilnet View in CoL

* Lophoziopsis propagulifera (Gottsche) Konstant. et Vilnet. View in CoL SVALBARD: Hodgetts 2015, Stotler & Crandall-Stotler 2017, Ellis et al. 2019b. GUSTAV V LAND: Konstantinova & Savchenko 2012. NOTE: The Southern Hemisphere Lophozia propagulifera has been synonymized with Lophoziopsis latifolia (= Lophoziopsis jurensis ) for some time, but this is rejected by Hodgetts et al. (2020).

Mannia Corda View in CoL

*** Mannia pilosa (Hornem.) Frye et L.Clark View in CoL [ Duvalia pilosa (Hornem.) Lindb. , Grimaldia pilosa (Hornem.) Lindb. , Neesiella pilosa (Hornem.) Schiffn. ]. SVALBARD: Kaalaas 1893a, Schiffner 1893, Bryhn 1906, Evans 1911, Jørgensen 1934, Müller 1952, Düll 1983 [with a ‘?’], Schuster 1992. GUSTAV V LAND: Berggren 1875a. SPITSBERGEN: Müller 1907b [Nordfjorden]. WEDEL JARLSBERG LAND: Święs & Karczmarz 1993b. NOTE: Mannia pilosa View in CoL was rejected from Svalbard by Frisvoll & Elvebakk (1996).

Marsupella Dumort. View in CoL

*** Marsupella sphacelata (Giesecke ex Lindenb.) Dumort. SVALBARD: Mohr 1901. NOTE: The mention in a list of Alabama bryophytes is certainly an error.

Mesoptychia (Lindb.) A.Evans

*** Mesoptychia bantriensis (Hook.) L.Söderstr. et Váňa View in CoL [ Lophozia bantriensis (Hook.) Steph. , Leiocolea bantriensis (Hook.) Jørg. View in CoL ]. SVALBARD: Müller 1954b, Arnell 1956, Schuster 1969, Schuster 1981, Smith 1990. NOTE: There seems not to be any primary report and Frisvoll & Elvebakk (1996) rejected it.

Mylia Gray View in CoL

*** Mylia anomala (Hook.) Gray. View in CoL SVALBARD: Müller 1956a, Smith 1990. NOTE: There seems not to exist any primary report and Frisvoll & Elvebakk (1996) rejected the occurrence. Nardia Gray View in CoL

*** Nardia breidleri (Limpr.) Lindb. View in CoL SVALBARD: Konstantinova 2000, Söderström et al. 2002 a, Hodgetts 2015. NOTE: Nardia breidleri View in CoL is only reported from Svalbard on a map without supporting data, but this is an error for Jan Mayen.

*** Nardia compressa (Hook.) Gray. View in CoL NOTE: Nardia compressa View in CoL was rejected from Svalbard by Hodgetts (2015; also Hodgetts & Lockhardt 2020) but we are not aware of any reports at all.

*** Nardia scalaris Gray View in CoL [ Jungermannia scalaris (Gray) Hook. , Alicularia scalaris (Gray) Corda View in CoL ]. SVALBARD: Müller 1909c, Damsholt 2002, 2013, Hodgetts 2015. GUSTAV V LAND: Hooker 1828. BÜNSOW LAND: Acock 1940. NOTE: Old reports are unreliable and it was rejected by Frisvoll & Elvebakk (1996).

Neoorthocaulis L.Söderstr., De Roo et Hedd. View in CoL

*** Neoorthocaulis attenuatus (Mårt.) L.Söderstr., De Roo et Hedd. View in CoL [ Jungermannia attenuata (Mårt.) Lindenb. , Lophozia attenuata (Mårt.) Dumort. ]. SVALBARD: Berggren 1875 b, Kaalaas 1893c as Jungermannia gracilis, Harmsen & Seidenfaden 1932 , Frye & Clark 1945 as Orthocaulis gracilis, Schuster 1969 , Worley 1972. BJØRNØYA: Berggren 1875a. GUSTAV V LAND: Wulff 1902 as Jungermannia gracilis . KONG KARLS LAND: Arnell 1900 as Jungermannia gracilis . ALBERT I LAND: Berggren 1875a. NORDENSKIÖLD LAND: Hagen 1952. PRINS KARLS FORLAND: Watson 1922, Summerhayes & Elton 1923a, 1923b. NOTE: A mainly boreal species and the many reports from Svalbard were all rejected by Frisvoll & Elvebakk (1996). The varieties described from Svalbard must belong to other species.

*** Neoorthocaulis binsteadii (Kaal.) L.Söderstr., De Roo et Hedd. View in CoL [ Barbilophozia binsteadii (Kaal.) Loeske , Lophozia binsteadii (Kaal.) A.Evans , Orthocaulis binsteadii (Kaal.) H.Buch ]. SVALBARD: Damsholt 2002, Söderström et al. 2002a, Konstantinova 2003, Ellis et al. 2012, Damsholt 2013, Hodgetts 2015. EDGEØYA: Heinemeijer 1979 as Orthocaulis cf. binsteadii . WEDEL JARLSBERG LAND: Konstantinova & Koroleva 2003. NOTE: Frisvoll & Elvebakk (1996) reject- ed it as all specimens they had seen could be referred to other species, mostly Barbilophozia hatcheri View in CoL .

*** Neoorthocaulis floerkei (F.Weber et D.Mohr) L.Söderstr., De Roo et Hedd. View in CoL [ Jungermannia floerkei F.Weber et D.Mohr , Barbilophozia floerkei (F.Weber et D.Mohr) Loeske , Lophozia floerkei (F.Weber et D.Mohr) Schiffn. , Orthocaulis floerkei (F.Weber et D.Mohr) H.Buch ]. SVALBARD: Berggren 1875b, Bryhn 1906, Jørgensen 1934, Frye & Clark 1945, Gradstein & Váňa 1987, Ellis et al. 2017. BJØRNØYA: Watson 1922. GUSTAV V LAND: Berggren 1875a. KONG KARLS LAND: Arnell 1900. ALBERT I LAND: Berggren 1875a, Arnell & Mårtensson 1959. HAAKON VII LAND: Arnell & Mårtensson 1959. NORDENSKIÖLD LAND: Ellis et al. 2016b. Olav V Land: Berggren 1875a. OSCAR II LAND: Gugnacka-Fiedor & Nryśkiewicz 1982, Boińska & Gugnacka-Fiedor 1983, Virtanen et al. 2013. PRINS KARLS FORLAND: Arnell 1900, Hagen 1908. NOTE: All reports of were rejected as other species by Frisvoll & Elvebakk (1996), many as Barbilophozia hatcheri View in CoL and Tritomaria quinquedentata The reports from Nordenskiöld Land and Oscar II Land are here rejected as Neoorthocaulis hyperboreus View in CoL (re-identified by N. Konstantinova).

Obtusifolium S.W.Arnell View in CoL

*** Obtusifolium obtusum (Lindb.) S.W.Arnell View in CoL [ Lophozia obtusa (Lindb.) A.Evans ]. SVALBARD: Bisang 1991, Söderström 1995, Bisang 1999, Paton 1999, Damsholt 2002. NOTE: There seems not to exist any primary report and the report was considered an error by Frisvoll & Elvebakk (1996). Odontoschisma (Dumort.) Dumort. View in CoL

*** Odontoschisma denudatum (Mårt.) Dumort. WEDEL JARLSBERG LAND View in CoL : Święs & Karczmarz 1991b. NOTE: Rejected by Frisvoll & Elvebakk (1996).

Checklist of liverworts on Svalbard 385

*** Odontoschisma sphagni (Dicks.) Dumort. OSCAR II LAND : Summerhayes & Elton 1923a, 1923b, Ellis et al. 2016a. NOTE: Rejected by Frisvoll & Elvebakk (1996). The report in Ellis et al. (2016a) is also certainly wrong.

Orthocaulis H.Buch

* Orthocaulis cavifolius H.Buch et S.W.Arnell View in CoL [ Sphenolobus cavifolius (H.Buch et S.W.Arnell) Müll.Frib. ]. SVALBARD: Düll 1983 [with a ‘?’], Konstantinova & Savchenko 2020b. HEER LAND: Philippi 1973. NOTE: Rejected as Sphenolobus minutus View in CoL by Frisvoll & Elvebakk (1996).

Pellia Raddi View in CoL

*** Pellia neesiana (Gottsche) Limpr. SVALBARD: Lindberg & Arnell 1889 as Marsilea neesii . NOTE: This old report must be rejected.

Plagiochila (Dumort.) Dumort.

*** Plagiochila porelloides (Torr. ex Nees) Lindenb. View in CoL SVALBARD: Düll 1983. NOTE Rejected as an error by Frisvoll & Elvebakk (1996).

*** Plagiochila spinulosa (Dicks.) Dumort. View in CoL SVALBARD: Phips 1774 as Lichenastrum ramosius , foliis trifidis. NOTE: Only mentioned through a doubtful indirect synonym and rejected by Frisvoll & Elvebakk (1996).

Pleurozia Dumort.

*** Pleurozia purpurea Lindb. NOTE: Rejected from Svalbard by Söderström et al. (2007) as it has only been reported from Jan Mayen, and the occurrence there is also highly questionable.

Protolophozia (R.M.Schust.) Schljakov

*** Protolophozia elongata (Steph.) Schljakov View in CoL [ Lophozia elongata Steph. , Orthocaulis elongatus (Steph.) A.Evans ]. SVALBARD: Herzog 1926, Ladyzhenskaja & Zhukova 1972, Lammes 1977, Düll 1983, Hodgetts 2015. NORDENSKIÖLD LAND: Arnell & Mårtensson 1959. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967. NOTE: Rejected by Frisvoll & Elvebakk (1996).

Ptilidium Nees View in CoL

*** Ptilidium pulcherrimum (Weber) Vain. View in CoL SVALBARD: Frye & Clark 1943, Söderström 1995. HAAKON VII LAND: Polunin 1945. NORDENSKIÖLD LAND: Hagen 1952. PRINS KARLS FORLAND: Watson 1922. NOTE: Rejected by Frisvoll & Elvebakk (1996).

Riccia View in CoL L.

*** Riccia sorocarpa Bisch. View in CoL SVALBARD: Müller 1952, Smith 1990, Damsholt 2002 [with a “?”], Damsholt 2013. NOTE: It is unclear what Müller based his report on and Söderström et al. (2007) rejected it.

Scapania (Dumort.) Dumort. View in CoL

*** Scapania aspera M.Bernet et Bernet. SVALBARD: Smith 1990. NOTE: We do not know what Smith based his reference on, but Scapania aspera do not occur on Svalbard.

*** Scapania calcicola (Arnell et J.Perss.) Ingham. View in CoL SVALBARD: Schuster 1974, Schljakov 1981, Düll 1983 [with a ‘?’], Smith 1990, Söderström 1995, Paton 1999, Damsholt 2002, Söderström et al. 2002a, Söderström et al. 2002b [with a ‘?’], Hodgetts 2015, Hodgetts & Lockhart 2020, Unan et al. 2021. DICKSON LAND: Konstantinova & Savchenko 2008c. HAAKON VII LAND: Arnell & Mårtensson 1959, Frisvoll & Elvebakk 1996. JAMES LAND: Frisvoll & Blom 1992. OSCAR II LAND: Arnell & Mårtensson 1959, Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997. NOTE: All reports are s.lat. (including “subsp. ligulifolia View in CoL ”) and is here rejected as Scapania ligulifolia View in CoL as only the latter is reported s.str. from the Arctic.

*** Scapania helvetica Gottsche. OSCAR II LAND View in CoL : Watson 1922 as Scapania curta var. geniculata . NOTE: Rejected by Frisvoll & Elvebakk (1996).

*** Scapania lingulata H.Buch. NOTE: Rejected from Svalbard by Söderström et al. (2002) but there seems not to be any earlier report at all to reject.

*** Scapania mucronata H.Buch. View in CoL SVALBARD: Düll 1983, Söderström 1995, Frisvoll & Elvebakk 1996, Söderström et al. 2002a, Konstantinova 2003, Hodgetts 2015, Hodgetts & Lockhart 2020. NORDENSKIÖLD LAND: Hagen 1952, Hadač 1989, Frisvoll & Blom 1997. OSCAR II LAND: Frisvoll & Blom 1997. NOTE: All reports are s. lat. or with doubt and we assume all refer to Scapania praetervisa View in CoL or the very variable Scapania obcordata View in CoL .

*** Scapania nemorea View in CoL (L.) Grolle. SVALBARD: Berggren 1875b as Scapania nemorosa . ALBERT I LAND: Livesay 1870 as Jungermannia nemorosa, Berggren 1875a as Scapania nemorosa . NOTE: The old reports are rejected by Frisvoll & Elvebakk (1996).

*** Scapania scandica (Arnell et H.Buch) Macvicar. View in CoL SVALBARD: Schljakov 1981, Söderström 1995, Damsholt 2002, Konstantinova 2003. WEDEL JARLSBERG LAND: Święs & Karczmarz 1991a. NOTE: Only reported s.lat. and we think all belong to Scapania parvifolia View in CoL .

*** Scapania undulata View in CoL (L.) Dumort. SVALBARD: Berggren 1875b, Müller 1901, 1915a, Bryhn 1906, Jørgensen 1934, Smith 1990. BJØRNØYA: Berggren 1875a. GUSTAV V LAND: Berggren 1875a as Scapania nemorosa var. purpurascens . ALBERT I LAND: Berggren 1875a also as Scapania undulata var. integrifolia and Scapania nemorosa var. purpurascens . OSCAR II LAND: Newsham & Goodall-Copestake 2021. NOTE: Rejected as partly Scapania tundrae View in CoL and partly Scapania hyperborea View in CoL by Frisvoll & Elvebakk (1996).

Solenostoma Mitt. View in CoL

*** Solenostoma gracillimum (Sm.) R.M.Schust. View in CoL [ Jungermannia gracillima Sm. ]. SVALBARD: Buch & Fagerström [rejecting earlier reports], Váňa 1974. NORDENSKIÖLD LAND: Berggren 1875a as Jungermannia genthiana . WEDEL JARLSBERG LAND: Święs & Karczmarz 1991b as Solenostoma crenulatum var. nana . NOTE: Rejected from Svalbard already by Buch & Fagerström (1946; see also Frisvoll & Elvebakk, 1976).