Soderstrom, , N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT, N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT

372 L SÖDERSTRÖM, N.A. KONSTANTINOVA, A. HAGBORG, T. PRESTØ, K. HASSEL & M. VON KONRAT

This publication. WEDEL JARLSBERG LAND: Eurola 1968. NOTE: Frisvoll & Elvebakk (1996) stated that the presence of Lophoziopsis excisa on Svalbard “needs confirmation”. However, more recent works have shown that the species occurs on Svalbard, but often it is scattered among other bryophytes, never occurring in large mats and therefore, perhaps, overlooked. Only reported with doubt from Bjørnøya and Prins Karls Forland.

** — var. elegans (R.M.Schust.) Konstant. et Vilnet [ Lophozia excisa var. elegans R.M.Schust. ]. SVALBARD: Hodgetts 2015, Hodgetts & Lockhart 2020. BARENTSØYA: Konstantinova & Savchenko 2018. HAAKON VII LAND: Konstantinova & Savchenko 2006, 2008a.

** — var. excisa . SVALBARD: Schljakov 1980 as Lophozia major, Söderström et al. 2007 , Hodgetts 2015, Hodgetts & Lockhart 2020.? BJØRNØYA: Watson 1922 as Lophozia excisa var. cylindracea with doubt. BARENTSØYA: Konstantinova & Savchenko 2018.? PRINS KARLS FORLAND: Watson 1922 as Lophozia excisa var. cylindracea with doubt.

** Lophoziopsis jurensis (Meyl. ex Müll.Frib.) Mamontov et Vilnet. SVALBARD: Schuster 1969 as Lophozia latifolia, Schuster & Damsholt 1974 as Lophozia latifolia, Steere & Inoue 1978 as Lophozia latifolia, Schljakov 1980 as Lophozia jurensis, Düll 1983 as Lophozia latifolia, Frisvoll & Blom 1992 as Lophozia latifolia, Söderström 1995 as Lophozia latifolia, Frisvoll & Elvebakk 1996 as Lophozia latifolia [“in need of confirmation”], Grolle & Long 2000 as Lophozia latifolia, Damsholt 2002 as Lophozia latifolia, Söderström et al. 2002a as Lophozia latifolia, Söderström et al. 2002b as Lophozia latifolia, Damsholt 2013 as Lophozia latifolia, Hodgetts & Lockhart 2020 . GUSTAV V LAND: Konstantinova & Savchenko 2012 as Lophoziopsis propagulifera View in CoL . PRINS OSCARS LAND: This publication. BARENTSØYA: This publication. (ALBERT I LAND: Arnell & Mårtensson 1959 as Lophozia latifolia [rejected by Grolle 1967]). (HAAKON VII LAND: Arnell & Mårtensson 1959 as Lophozia latifolia [rejected by Grolle 1967]). (NORDENSKIÖLD LAND: Arnell & Mårtensson 1959 as Lophozia latifolia [rejected by Grolle 1967], Eurola 1971 as Lophozia cf. latifolia ). OSCAR II LAND: Arnell & Mårtensson 1959 as Lophozia latifolia, Grolle 1967 as Lophozia latifolia, Frisvoll & Blom 1997 as Lophozia latifolia . NOTE: All reports of Lophozia latifolia (= Lophoziopsis jurensis ) from Svalbard, except one, were rejected by Grolle (1967), but later reports confirm its presence there.

*** Lophoziopsis longidens (Lindb.) Konstant. et Vilnet View in CoL [ Jungermannia longidens Lindb. , Lophozia longidens (Lindb.) Macoun ]. SVALBARD: Arnell 1928, Jørgensen 1934, Frye & Clark 1945, Müller 1954b, Arnell 1956, Smith 1990, Söderström 1995, Paton 1999, Bapna & Kachroo 2000b, Damsholt 2002, Söderström et al. 2002a, 2002b, Damsholt 2013. BJØRNØYA: Watson 1922, Summerhayes & Elton 1923a, Söderström 1995, Frisvoll & Elvebakk 1996 Damsholt 2002, Söderström et al. 2002b. KONG KARLS LAND: Arnell 1900, Müller 1910b, Frisvoll & Elvebakk 1996. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2008b. OSCAR II LAND: Watson 1922, Summerhayes & Elton 1923a, 1923b, Frisvoll & Elvebakk 1996. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Karczmarz & Święs 1989, Frisvoll & Elvebakk 1996.

** — subsp. longidens . SVALBARD: Schuster 1969 [with doubt], Düll 1983 [with a ‘?’], Söderström et al. 2002b, Hodgetts 2015, Hodgetts & Lockhart 2020. BJØRNØYA: Söderström et al. 2002b.

*** Lophoziopsis pellucida (R.M.Schust.) Konstant. et Vilnet View in CoL [ Lophozia pellucida R.M.Schust. ]. SVALBARD: Söderström 1995, Grolle & Long 2000, Konstantinova 2000, Damsholt 2002, Söderström et al. 2002b, Hodgetts 2015, Konstantinova & Savchenko 2020b. NORDAUSTLANDET: Konstantinova & Savchenko 2015. DICKSON LAND: This publication. HAAKON VII LAND: Frisvoll & Blom 1992, 1997, Frisvoll & Elvebakk 1996. NORDENSKIÖLD LAND: Frisvoll & Blom 1997. NY-FRIESLAND: This publication.

** — var. minor (R.M.Schust.) L.Söderstr. et Váňa [ Lophozia pellucida var. minor R.M.Schust. ]. SVALBARD: Söderström et al. 2002a, Söderström et al. 2007, Hodgetts & Lockhart 2020. GUSTAV V LAND: Konstantinova & Savchenko 2012. SPITSBERGEN: Bakalin 2005 [W Svalbard Isl.]. NORDENSKIÖLD LAND: Schuster & Konstantinova 1996.

** — var. pellucida . SVALBARD: Söderström et al. 2002a, Söderström et al. 2007, Hodgetts & Lockhart 2020. GUSTAV V LAND: Konstantinova & Savchenko 2012. DICKSON LAND: Bakalin 2005.

*** Lophoziopsis polaris (R.M.Schust.) Konstant. et Vilnet View in CoL [ Lophozia polaris (R.M.Schust.) R.M.Schust. et Damsh. ]. SVALBARD: Düll 1983 [with a ‘?’], Söderström 1995, Konstantinova 2000, Damsholt 2002, Söderström et al. 2002a, 2002b, Vilnet et al. 2010, Konstantinova et al. 2020a. GUSTAV V LAND: Konstantinova & Savchenko 2012, 2015. PRINS OSCARS LAND: Konstantinova & Savchenko 2015, 2020b. BARENTSØYA: Konstantinova et al. 2014, Konstantinova & Savchenko 2018. DICKSON LAND: This publication. HAAKON VII LAND: Konstantinova & Savchenko 2006. HEER LAND: Konstantinova et al. 2020b. JAMES LAND: This publication. NATHORST LAND: This publication. NORDENSKIÖLD LAND: Hadač 1989, Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997, Konstantinova et al. 2020b. NY-FRIESLAND: This publication. OSCAR II LAND: Koroleva & Borovichev 2014. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. SØRKAPP LAND: This publication. WEDEL JARLSBERG LAND: This publication.

** — var. polaris View in CoL . SVALBARD: Hodgetts 2015, Hodgetts & Lockhart 2020. PRINS OSCARS LAND: Konstantinova & Savchenko 2008a. HAAKON VII LAND: Konstantinova & Savchenko 2008a. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2008a.

** — var. sphagnorum (R.M.Schust.) Konstant. et Vilnet [ Lophozia polaris var. sphagnorum (R.M.Schust.) R.M.Schust. et Damsh. ]. SVALBARD: Hodgetts 2015, Hodgetts & Lockhart 2020. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2007, 2008 a, 2008b.

** Lophoziopsis rubrigemma (R.M.Schust.) Konstant. et Vilnet View in CoL [ Lophozia rubrigemma R.M.Schust. ]. SVALBARD: Konstantinova 2000, Söderström et al. 2002 a, Hodgetts 2015, Hodgetts & Lockhart 2020, Konstantinova & Savchenko 2020b. GUSTAV V LAND: Konstantinova & Savchenko 2012, 2015.? BARENTSØYA: Konstantinova & Savchenko 2018 as cf. rubrigemma .? NORDENSKIÖLD LAND: Konstantinova & Savchenko 2008b as cf. rubrigemma . NOTE: Lophoziopsis rubrigemma View in CoL has only been reported with some doubt from Barentsøya and Nordenskiöld Land.

Mannia Corda View in CoL

*** Mannia triandra (Scop.) Grolle. View in CoL SVALBARD: Borovichev et al. 2015, Hodgetts 2015 [with a ‘?’], Borovichev & Bakalin 2016, Hodgetts & Lockhart 2020, Konstantinova & Savchenko 2020b. OSCAR II LAND: Borovichev 2010.

Checklist of liverworts on Svalbard 373

Marchantia L.

*** Marchantia polymorpha View in CoL L. SVALBARD: Kaalaas 1893a, Bryhn 1906, Müller 1907b, Jørgensen 1934, Müller 1952, Bischler 1984, Bischler-Causse 1989, Smith 1990, Söderström et al. 2002 b, Konstantinova 2003. BJØRNØYA: Berggren 1875a, Söderström et al. 2002b. GUSTAV V LAND: Berggren 1875a, 1875b. KONG KARLS LAND: Arnell 1900. SPITSBERGEN: Sommerfeldt 1833 [Vest-Spitsbergen], Polunin 1945 [Kongsfjorden]. ALBERT I LAND: Berggren 1875a. NORDENSKIÖLD LAND: Hadač 1946, 1989. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Karczmarz & Święs 1989, Święs & Karczmarz 1991b.

*** — subsp. montivagans Bischl. et Boissel.-Dub. View in CoL SVALBARD: Düll 1983 as Marchantia alpestris, Söderström 1995 View in CoL as Marchantia alpestris, Frisvoll & Elvebakk 1996 View in CoL as Marchantia alpestris, Paton 1999 View in CoL , Damsholt 2002 as Marchantia alpestris, Söderström et al. 2002a View in CoL , Söderström et al. 2002b, Damsholt 2013 as Marchantia alpestris, Hodgetts 2015 View in CoL , Hodgetts & Lockhart 2020. BJØRNØYA: Söderström 1995 as Marchantia alpestris, Frisvoll & Elvebakk 1996 View in CoL as Marchantia alpestris, Damsholt 2002 View in CoL as Marchantia alpestris, Söderström et al. 2002b View in CoL . GUSTAV V LAND: Lindberg 1866 as Marchantia polymorpha View in CoL B alpestris, Konstantinova & Savchenko 2012 View in CoL . BARENTSØYA: Philippi 1973 as Marchantia alpestris, Konstantinova & Savchenko 2018 View in CoL . EDGEØYA: Philippi 1973 as Marchantia alpestris View in CoL . ALBERT I LAND: Lindberg 1866 as Marchantia polymorpha View in CoL B alpestris View in CoL . DICKSON LAND: Belkina et al. 2015. HAAKON VII LAND: Arnell & Mårtensson 1959 as Marchantia alpestris View in CoL . JAMES LAND: This publication. NORDENSKIÖLD LAND: Arnell & Mårtensson 1959 as Marchantia alpestris, Konstantinova & Savchenko 2008b View in CoL as Marchantia alpestris, Prestø et al. 2014 View in CoL . OSCAR II LAND: Arnell & Mårtensson 1959 as Marchantia alpestris, Prestø et al. 2014 View in CoL . PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967 as Marchantia alpestris View in CoL . NOTE: We are not aware of any primary report of subsp. montivagans View in CoL from Bjørnøya and Dickson Land.

*** — subsp. ruderalis Bischl. et Boissel.-Dub. SVALBARD: Düll 1983 as, Söderström 1995 as Marchantia polymorpha, Frisvoll & Elvebakk 1996 as Marchantia polymorpha, Damsholt 2002 as Marchantia latifolia, Söderström et al. 2002a , Söderström et al. 2002b, Hodgetts 2015, Ellis et al. 2019 a, Hodgetts & Lockhart 2020. DICKSON LAND: This publication. WEDEL JARLSBERG LAND: Frisvoll & Blom 1997 as Marchantia polymorpha . NOTE: Subsp. ruderalis has only been confirmed from anthropogenic areas and all other reports should probably be subsp. montivagans .

*** Marchantia quadrata Scop. View in CoL [ Preissia quadrata (Scop.) Nees , Chomiocarpon quadratum (Scop.) Lindb. View in CoL ]. SVALBARD: Schiffner 1893, Bryhn 1906 as Chomocarpon commutatus, Müller 1907b , Jørgensen 1934, Horikawa 1950 as Preissia quadrata, Düll 1983 , Smith 1990, Söderström 1995, Frisvoll & Elvebakk 1996, Paton 1999, Damsholt 2002, Söderström et al. 2002b, Long & Crandall-Stotler 2020. BJØRNØYA: Watson 1922, Summerhayes & Elton 1923a, Söderström 1995, Frisvoll & Elvebakk 1996, Damsholt 2002, Söderström et al. 2002b. GUSTAV V LAND: Konstantinova & Savchenko 2012. PRINS OSCARS LAND: This publication. BARENTSØYA: Philippi 1973, Konstantinova & Savchenko 2018. DICKSON LAND: Belkina et al. 2015. HAAKON VII LAND: Arnell & Mårtensson 1959, Frisvoll 1978. NATHORST LAND: This publication. NORDENSKIÖLD LAND: This publication, Prestø et al. 2014. OSCAR II LAND: Arnell & Mårtensson 1959. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967.

* — subsp. hyperborea (R.M.Schust.) Borovich. [ Preissia quadrata subsp. hyperborea R.M.Schust. ]. SVALBARD: Söderström et al. 2007, Hodgetts 2015, Hodgetts & Lockhart 2020. BARENTSØYA: Konstantinova & Savchenko 2018. SPITSBERGEN: Thingsgaard & Damsholt 2007. DICKSON LAND: This publication. HAAKON VII LAND: This publication. JAMES LAND: This publication. OSCAR II LAND: This publication. PRINS KARLS FORLAND: This publication.

* — subsp. quadrata . SVALBARD: Schuster 1992, Söderström et al. 2002a, Damsholt 2013, Hodgetts 2015, Hodgetts & Lockhart 2020. NOTE: It is possible that only subsp. hyperborea occurs on Svalbard.

*** Marchantia romanica (Radian) D.G.Long, Crand.-Stotl., L.L.Forrest et J.C.Villarreal View in CoL [ Bucegia romanica Radian View in CoL ]. SVALBARD: Ellis et al. 2016a, Xiang et al. 2016, Sofronova & Potemkin 2018, Hodgetts & Lockhart 2020. NORDAUSTLANDET: Xiang et al. 2016. GUSTAV V LAND: Konstantinova et al. 2014. PRINS OSCARS LAND: Konstantinova et al. 2014. BARENTSØYA: Konstantinova et al. 2014 Konstantinova & Savchenko 2018. DICKSON LAND: Konstantinova et al. 2014. HAAKON VII LAND: This publication. OSCAR II LAND: This publication. SABINE LAND: Konstantinova et al. 2014.

Marsupella Dumort. View in CoL

*** Marsupella apiculata Schiffn. View in CoL [ Gymnomitrion apiculatum (Schiffn.) Müll.Frib. View in CoL ]. SVALBARD: Schuster & Damsholt 1974, Schuster 1974, Godfrey & Schofield 1979, Schljakov 1981, Düll 1983, Söderström 1995, Paton 1999, Damsholt 2002, Söderström et al. 2002a, 2002 b, Konstantinova 2003, Vilnet et al. 2010, Hodgetts 2015, Mamontov et al. 2018, Mamontov et al. 2019, Hodgetts & Lockhart 2020. PRINS OSCARS LAND: Konstantinova & Savchenko 2008 a, Konstantinova & Savchenko 2015. ALBERT I LAND: Arnell & Mårtensson 1959, Frisvoll & Blom 1992, Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997, Thingsgaard & Damsholt 2007, Konstantinova & Savchenko 2008a. HAAKON VII LAND: Frisvoll & Blom 1992, Frisvoll & Blom 1997. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2007, 2008 a, 2008b, Konstantinova et al. 2020b. OSCAR II LAND: Kobayashi et al. 1990, Frisvoll & Blom 1992, 1997, Frisvoll & Elvebakk 1996.

*** Marsupella arctica (Berggr.) Bryhn et Kaal. View in CoL [ Sarcoscyphos emarginatus var. arcticus Berggr. , Marsupella emarginata var. arctica (Berggr.) Frye et L.Clark View in CoL ]. SVALBARD: Berggren 1875b, Bryhn 1906, Herzog 1926 as Marsupella groenlandica, Schuster 1950 View in CoL , Müller 1956a, Persson & Gjaerevoll 1961, Schuster & Damsholt 1974, Schljakov 1981, Düll 1983, Geissler & Bischler 1985, Geissler & Bischler 1985, Vitt et al. 1987, Long et al. 1990, Söderström 1995, Schuster & Konstantinova 1996, Paton 1999, Konstantinova 2000, Söderström et al. 2002a, 2002b, Vilnet et al. 2010, Hodgetts 2015, Shaw et al. 2015, Mamontov et al. 2018, 2019, Hodgetts & Lockhart 2020. NORDAUSTLANDET: Konstantinova & Savchenko 2015, 2020b. PRINS OSCARS LAND: Konstantinova & Savchenko 2008 a, Konstantinova et al. 2020b. SPITSBERGEN: Vilnet et al. 2011. ALBERT I LAND: TYPE, Berggren 1875a, Müller 1909a, Frye & Clark 1943, Schuster 1974, Grolle 1976, Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997, Damsholt 2002, 2013. NORDENSKIÖLD LAND: