Soderstrom, , N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT, N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT

370 L SÖDERSTRÖM, N.A. KONSTANTINOVA, A. HAGBORG, T. PRESTØ, K. HASSEL & M. VON KONRAT

Harmsen & Seidenfaden 1932, Jørgensen 1934, Frye & Clark 1943, Steere 1954, Arnell 1956, Müller 1956a, Worley 1972, Schuster 1974, Schljakov 1981, Smith 1990, Söderström 1995, Frisvoll & Elvebakk 1996, Paton 1999, Damsholt 2002, Söderström et al. 2002a, 2002b, Hodgetts 2015, Shaw et al. 2015, Patzak et al. 2016 a, Hodgetts & Lockhart 2020. BJØRNØYA: Berggren 1875a, Söderström 1995, Frisvoll & Elvebakk 1996, Damsholt 2002, Söderström et al. 2002b. GUSTAV V LAND: Lindberg 1866, Berggren 1875a. PRINS OSCARS LAND: Konstantinova & Savchenko 2008a. BARENTSØYA: Philippi 1973, Konstantinova & Savchenko 2018. EDGEØYA: Philippi 1973. ALBERT I LAND: Lindberg 1866, Berggren 1875a, Arnell & Mårtensson 1959, Thingsgaard & Damsholt 2007, Konstantinova & Savchenko 2008a. DICKSON LAND: This publication, Belkina et al. 2015. HAAKON VII LAND: Arnell & Mårtensson 1959. NORDENSKIÖLD LAND: Berggren 1875 a, Hagen 1952, Eurola 1968, Hadač 1989, Konstantinova & Savchenko 2007, 2008 a, 2008b, Virtanen et al. 2013. OSCAR II LAND: Arnell & Mårtensson 1959. PRINS KARLS FORLAND: Berggren 1875 a, Konstantinova & Savchenko 2020a. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Karczmarz & Święs 1988, 1990b, Święs & Karczmarz 1991a, 1991b, 1993a.

*** Gymnomitrion corallioides Nees View in CoL [ Cesius corallioides (Nees) Carruth. View in CoL ‘ Cesia corallioides ’]. SVALBARD: Berggren 1875b, Bryhn 1906, Müller 1909a, Jørgensen 1934, Frye & Clark 1943, Steere 1954, Schuster & Damsholt 1974, Schuster 1974, Düll 1983, Smith 1990, Söderström 1995, Frisvoll & Elvebakk 1996, Paton 1999, Damsholt 2002, Söderström et al. 2002 a, Hentschel et al. 2006, 2007, Heinrichs et al. 2007, Vilnet et al. 2010, Damsholt 2013, Hodgetts 2015, Shaw et al. 2015, Mamontov et al. 2018, Mamontov et al. 2019, Hodgetts & Lockhart 2020. BJØRNØYA: Söderström 1995, Frisvoll & Elvebakk 1996, Damsholt 2002. GUSTAV V LAND: Berggren 1875a. PRINS OSCARS LAND: Konstantinova & Savchenko 2008a. BARENTSØYA: Hoffman 1968, Philippi 1973, Konstantinova & Savchenko 2018. EDGEØYA: Philippi 1973. KONG KARLS LAND: Arnell 1900, Hoffman 1968. SPITSBERGEN: Arnell & Mårtensson 1959 [W. Spitsbergen], Vilnet et al. 2011. ALBERT I LAND: Berggren 1875 a, Frisvoll 1978. DICKSON LAND: This publication. HEER LAND: Philippi 1973. NORDENSKIÖLD LAND: Hadač 1946, Hagen 1952, Eurola 1968, Hadač 1989, Konstantinova & Savchenko 2007, 2008 a, 2008b, Thingsgaard & Damsholt 2007, Virtanen et al. 2013, Konstantinova et al. 2020b. NY-FRIESLAND: This publication. OSCAR II LAND: This publication. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. SABINE LAND: Philippi 1973. SØRKAPP LAND: Dubiel & Olech 1990, Wietrzyk et al. 2016. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Eurola 1968, Bednarek-Ochyra et al. 1987b, Karczmarz & Święs 1990a. NOTE: We are not aware of any primary report from Bjørnøya.

* — var. minutum Lindb. ALBERT I LAND : TYPE, Lindberg 1866. HAAKON VII LAND: Lindberg 1866. NOTE: The variety is not recognized in recent treatments, and we do not think it is worth to recognize either. However, we do not know if this name refers to Gymnomitrion coralloides or to any other species as we have not seen any type material.

Haplomitrium Nees

*** Haplomitrium hookeri (Lyell ex Sm.) Nees. View in CoL SVALBARD: Grolle 1964, Schuster 1966, 1967, Worley 1969, Schofield & Crum 1972, Schuster & Damsholt 1974, Schljakov 1976, Godfrey 1977, Engel 1981, Düll 1983, Schuster 1983, Smith 1990, Bartholomew-Began 1991, Parihar et al. 1994, Söderström 1995, Schuster & Konstantinova 1996, Paton 1999, Schumacker et al. 1999, Bapna & Kachroo 2000 a, Konstantinova 2000, 2003, Damsholt 2002, Söderström et al. 2002a, 2002b, Sotiaux et al. 2006, Engel & Glenny 2008, Damsholt 2013, Duckett & Slack 2013, Hodgetts 2015, Hodgetts & Lockhart 2020. NORDENSKIÖLD LAND: Arnell & Mårtensson 1959, Frisvoll & Blom 1992, Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997.

Isopaches H.Buch View in CoL

*** Isopaches bicrenatus (Schmidel ex Hoffm.) H.Buch View in CoL [ Lophozia bicrenata (Schmidel ex Hoffm.) Dumort. ]. SVALBARD: Frye & Clark 1945, Schuster 1969 [‘not confirmed’], Düll 1983, Söderström 1995, Paton 1999, Damsholt 2002, 2013, Söderström et al. 2002a, 2002b, Hodgetts 2015, Hodgetts & Lockhart 2020. BÜNSOW LAND: Summerhayes & Elton 1923 a, Frisvoll & Blom 1997. NORDENSKIÖLD LAND: Watson 1922, Summerhayes & Elton 1923b, Konstantinova & Savchenko 2008b. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Frisvoll & Blom 1992, 1997, Frisvoll & Elvebakk 1996.

Jungermannia View in CoL L.

*** Jungermannia polaris Lindb. View in CoL [ Jungermannia pumila * polaris (Lindb.) Berggr. View in CoL , Aplozia polaris (Lindb.) Bryhn et Kaal. View in CoL ‘ Haplozia ’, Jungermannia pumila subsp. polaris (Lindb.) R.M.Schust. View in CoL , Solenostoma polare (Lindb.) R.M.Schust. ]. SVALBARD: Berggren 1875b, Bryhn 1906,, Herzog 1926, Arnell 1928, Schuster et al. 1959, Düll 1983, Smith 1990, Söderström 1995, Paton 1999, Söderström et al. 2002a, Söderström et al. 2002b, Hodgetts 2015, Konstantinova & Vilnet 2016, Hodgetts & Lockhart 2020. BJØRNØYA: Berggren 1875a, Váňa 1973, Söderström 1995, Frisvoll & Elvebakk 1996, Damsholt 2002, Söderström et al. 2002b. GUSTAV V LAND: Konstantinova & Savchenko 2012. PRINS OSCARS LAND: This publication. BARENTSØYA: Konstantinova & Savchenko 2018. DICKSON LAND: Frisvoll 1981, Konstantinova & Savchenko 2008 c, 2011. HAAKON VII LAND: Lindberg 1866, Arnell & Mårtensson 1959, Arnell & Mårtensson 1959 as Jungermannia schiffneri, Váňa 1973 View in CoL , Grolle 1976, Geissler & Bischler 1987, Konstantinova & Savchenko 2006. HEER LAND: This publication. JAMES LAND: This publication. NATHORST LAND: This publication. NORDENSKIÖLD LAND: Berggren 1875 a, Konstantinova & Savchenko 2007, 2008 a, 2008b. NY-FRIESLAND: TYPE, Schuster 1969, Frisvoll & Elvebakk 1996, Damsholt 2002, 2013. OSCAR II LAND: Arnell & Mårtensson 1959. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. SØRKAPP LAND: This publication. WEDEL JARLSBERG LAND: This publication.

*** Jungermannia pumila With. View in CoL [ Solenostoma pumilum (With.) Müll.Frib. ]. SVALBARD: Schuster 1969, Worley 1972, Váňa 1973, Söderström 1995, Damsholt 2002 [with a?], Konstantinova 2003, Damsholt 2013, Schäfer-Verwimp et al. 2019. BJØRNØYA: Berggren 1875a, Söderström 1995. DICKSON LAND: This publication. HAAKON VII LAND: Arnell & Mårtensson 1959. NATHORST LAND: This publication. OSCAR II LAND: Arnell & Mårtensson 1959. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. SØRKAPP LAND: This publication. WEDEL JARLSBERG LAND: This publication. NOTE: Many identifications are s.lat. or only preliminary and may belong to Jungermannia polaris View in CoL .

Checklist of liverworts on Svalbard 371

Lophozia (Dumort.) Dumort.

** Lophozia fuscovirens Bakalin et Vilnet. SVALBARD: Hodgetts & Lockhart 2020, Hodgetts et al. 2020, Konstantinova et al. 2020a. HEER LAND: This publication. PRINS KARLS FORLAND: Ellis et al. 2019c, Konstantinova & Savchenko 2020a.

** Lophozia murmanica Kaal. View in CoL SVALBARD: Herzog 1926 as Sphenolobus groenlandicus, Steere 1954 as Lophozia groenlandica, Arnell 1956 as Lophozia groenlandica, Schuster 1961 as Lophozia groenlandica, Schljakov 1980 as Lophozia groenlandica, Düll 1983 as Lophozia groenlandica, Damsholt 2002 , Konstantinova 2003 as Lophozia wenzelii var. groenlandica, Söderström et al. 2007 as Lophozia groenlandica, Kalinauskaitë & von Cräutlein 2010 , Hodgetts 2015 as Lophozia groenlandica, Hodgetts & Lockhart 2020 , Konstantinova & Savchenko 2020b. GUSTAV V LAND: Konstantinova & Savchenko 2012 as Lophozia wenzelii var. groenlandica . BARENTSØYA: Konstantinova & Savchenko 2018. SPITSBERGEN: Bakalin 2004 as Lophozia wenzelii var. groenlandica [Kongsfjorden], Bakalin 2005 as Lophozia wenzelii var. groenlandica [Kings Bay]. ALBERT I LAND: Thingsgaard & Damsholt 2007. NORDENSKIÖLD LAND: Bryhn 1909 as Lophozia groenlandica, Müller 1910a as Sphenolobus groenlandicus, Konstantinova & Savchenko 2008a as Lophozia wenzelii var. groenlandica, Konstantinova & Savchenko 2008b as Lophozia wenzelii var. groenlandica . PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967 as Lophozia groenlandica, Karczmarz & Święs 1988 as Lophozia groenlandica, Karczmarz & Święs 1989 , 1990b both as Lophozia groenlandica, Święs & Karczmarz 1991a , 1991b, 1993b all as Lophozia groenlandica . NOTE: The nomenclature in Lophozia View in CoL is confusing, and the same name has been used for different species (see Söderström et al., 2013). The name Lophozia groenlandica is used for at least four Lophozia View in CoL taxa, L. murmanica View in CoL , L. wenzelii View in CoL , L. schusterana and L. ventricosa View in CoL . We think that many of the Svalbard records can be referred to our Lophozia murmanica View in CoL concept.

** Lophozia savicziae Schljakov. View in CoL SVALBARD: Konstantinova 2000, Söderström et al. 2002 a, Bakalin 2005 [Storfjord], Hodgetts 2015, Hodgetts & Lockhart 2020, Konstantinova & Savchenko 2020b. Olav V Land: This publication.

*** Lophozia silvicola H.Buch View in CoL [ Lophozia ventricosa var. silvicola (H.Buch) E.W.Jones View in CoL ]. SVALBARD: Söderström et al. 2007, Ellis et al. 2014 a, Hodgetts 2015, Hodgetts & Lockhart 2020. BARENTSØYA: Konstantinova & Savchenko 2018. ALBERT I LAND: Thingsgaard & Damsholt 2007.

*** Lophozia silvicoloides N.Kitag. View in CoL SVALBARD: Vilnet et al. 2005 as Lophozia silvicoloides var. paroica, Söderström et al. 2007 , Vilnet et al. 2010, Bakalin 2011, Fedosov et al. 2015, Hodgetts 2015, Bakalin & Vilnet 2019, Hodgetts & Lockhart 2020, Hodgetts et al. 2020, Konstantinova & Savchenko 2020b, Konstantinova et al. 2020a. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2007, 2008 a, 2008b.

* Lophozia subapiculata R.M.Schust. et Damsh. View in CoL SVALBARD: Hodgetts & Lockhart 2020, Hodgetts et al. 2020, Konstantinova & Savchenko 2020b. BARENTSØYA: Konstantinova & Savchenko 2018.

** Lophozia svalbardensis Konstant., Vilnet et Mamontov. GUSTAV V LAND : TYPE, Konstantinova et al. 2020a.

*** Lophozia ventricosa (Dicks.) Dumort. View in CoL [ Jungermannia ventricosa Dicks. ]. SVALBARD: Bryhn 1906, Harmsen & Seidenfaden 1932, Frye & Clark 1945, Söderström 1995, Paton 1999, Söderström et al. 2002a, 2002b, Hodgetts 2015, Hodgetts & Lockhart 2020. BJØRNØYA: Watson 1922, Summerhayes & Elton 1923a, Söderström 1995, Frisvoll & Elvebakk 1996, Söderström et al. 2002b. Frisvoll & Elvebakk 1996. KONG KARLS LAND: Arnell 1900, Frisvoll & Elvebakk 1996. ALBERT I LAND: Lindberg 1866 as Jungermannia ventricosa D * gemmipara, Jungermannia ventricosa b laxa and Jungermannia ventricosa E * pulchella, Frisvoll & Elvebakk 1996. NORDENSKIÖLD LAND: Stebel et al. 2018. OSCAR II LAND: Watson 1922, Summerhayes & Elton 1923a, 1923b, Frisvoll & Elvebakk 1996. PRINS KARLS FORLAND: Arnell 1900, Watson 1922, Frisvoll & Elvebakk 1996.

* — var. longiflora (Nees) Macoun [ Lophozia longiflora (Nees) Schiffn. ]. SVALBARD: Frye & Clark 1945. BJØRNØYA: Watson 1922. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2008b. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a.

* — var. ventricosa . SVALBARD: Damsholt 2002, Söderström et al. 2007, Damsholt 2013. PRINS OSCARS LAND: This publication. ALBERT I LAND: Thingsgaard & Damsholt 2007.

* Lophozia wenzelii (Nees) Steph. View in CoL [ Jungermannia wenzelii Nees ]. SVALBARD: Bryhn 1906, Müller 1910b, Arnell 1928 as Jungermannia wenzelii, Jørgensen 1934 , Frye & Clark 1945, Müller 1954b, Worley 1972, Smith 1990, Söderström 1995, Paton 1999, Söderström et al. 2002a, 2002b, Damsholt 2013, Hodgetts 2015, Hodgetts & Lockhart 2020. GUSTAV V LAND: This publication. PRINS OSCARS LAND: This publication. BARENTSØYA: Philippi 1973. EDGEØYA: Philippi 1973. KONG KARLS LAND: Arnell 1900. HAAKON VII LAND: Arnell & Mårtensson 1959, Newsham & Goodall-Copestake 2021. HEER LAND: Philippi 1973. NORDENSKIÖLD LAND: Arnell & Mårtensson 1959, Hadač 1989, Frisvoll & Elvebakk 1996. OSCAR II LAND: Arnell & Mårtensson 1959, Kobayashi et al. 1990. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Frisvoll & Elvebakk 1996.

* — var. lapponica H.Buch et S.W.Arnell. SVALBARD: Bakalin 2004, 2005, Vilnet et al. 2005, 2010, Söderström et al. 2007, Damsholt 2013, Bakalin & Vilnet 2019, Konstantinova et al. 2020a. BARENTSØYA: Konstantinova & Savchenko 2018. ALBERT I LAND: Thingsgaard & Damsholt 2007. NORDENSKIÖLD LAND: Konstantinova & Koroleva 2003, Konstantinova & Savchenko 2007, 2008b, Konstantinova et al. 2020b. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a.

* — var. wenzelii . SVALBARD: Arnell 1956, Schuster 1969, Schuster & Damsholt 1974, Düll 1983, Damsholt 2002. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2008b. Lophoziopsis Konstant. et Vilnet

*** Lophoziopsis excisa (Dicks.) Konstant. et Vilnet View in CoL [ Lophozia excisa (Dicks.) Dumort. ]. SVALBARD: Düll 1983, Damsholt 2002 [with a “?”], Söderström et al. 2002a, Vilnet et al. 2005, Vilnet et al. 2010, 2012a, Damsholt 2013, Mamontov & Vilnet 2017, Bakalin & Vilnet 2019, Konstantinova et al. 2020a.? BJØRNØYA: Söderström 1995. PRINS OSCARS LAND: This publication. ALBERT I LAND: Thingsgaard & Damsholt 2007. HEER LAND: This publication. NORDENSKIÖLD LAND: Arnell & Mårtensson 1959 as Lophozia kiaerii, Konstantinova & Savchenko 2007 , 2008 a, 2008b. OSCAR II LAND: Arnell & Mårtensson 1959 as Lophozia kiaerii, Newsham & Goodall-Copestake 2021 . SØRKAPP LAND: