Soderstrom, , N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT, N. A. KONSTANTINOVA, A. HAGBORG, T. PRESTO, K. HASSEL & M. VON KONRAT

368 L SÖDERSTRÖM, N.A. KONSTANTINOVA, A. HAGBORG, T. PRESTØ, K. HASSEL & M. VON KONRAT

Watson 1922, Summerhayes & Elton 1923b. SØRKAPP LAND: Dubiel & Olech 1990. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Karczmarz & Święs 1989, 1990a, Święs & Karczmarz 1991a, 1991b, Frisvoll & Elvebakk 1996 [with doubt], Frisvoll & Blom 1997. NOTE: Cephalozia bicuspidata has often been reported s.lat. (i.e. including Cephalozia ambigua ) and several reports have been, or should be, rejected as the latter species.

Cephaloziella (Spruce) Schiffn.

*** Cephaloziella arctogena (R.M.Schust.) Konstant. View in CoL SVALBARD: Söderström et al. 2007, Hodgetts 2015, Hodgetts & Lockhart 2020, Konstantinova & Savchenko 2020b. HAAKON VII LAND: Thingsgaard & Damsholt 2007.

*** Cephaloziella integerrima (Lindb.) Warnst. View in CoL [ Dichiton integerrimum (Lindb.) H.Buch ]. SVALBARD: Düll 1983, Smith 1990, Söderström 1995 [rejecting earlier reports], Damsholt 2002 [with a “?”], Söderström et al. 2002a [rejecting earlier reports], Söderström et al. 2002b [rejecting earlier reports], Damsholt 2013, Hodgetts 2015, Hodgetts & Lockhart 2020. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2007, 2008 a, 2008b, 2020b. OSCAR II LAND: Arnell & Mårtensson 1959, Frisvoll & Elvebakk 1996 [rejecting earlier reports], Konstantinova & Savchenko 2020b. SØRKAPP LAND: This publication. WEDEL JARLSBERG LAND: This publication. *** Cephaloziella polystratosa (R.M.Schust. et Damsh.) Konstant. View in CoL SVALBARD: Hodgetts 2015, Hodgetts & Lockhart 2020, Hodgetts et al. 2020. NORDAUSTLANDET: Konstantinova & Savchenko 2020b. GUSTAV V LAND: Konstantinova & Savchenko 2012, 2015.

*** Cephaloziella uncinata R.M.Schust. View in CoL SVALBARD: Schuster 1980, Düll 1983, Davison 1993, Söderström 1995, Grolle & Long 2000, Konstantinova 2000, Söderström et al. 2002a, 2002b, Prey et al. 2014, Hodgetts 2015, Hodgetts & Lockhart 2020. PRINS OSCARS LAND: Konstantinova & Savchenko 2008a. ALBERT I LAND: Thingsgaard & Damsholt 2007, Konstantinova & Savchenko 2008a. DICKSON LAND: This publication, Frisvoll & Elvebakk 1996. HAAKON VII LAND: This publication. NORDENSKIÖLD LAND: Hadač 1946 as Cephalozia striatula [cf. Frisvoll & Elvebakk 1996], Frisvoll & Elvebakk 1996, Damsholt 2002, Konstantinova & Savchenko 2008b, Damsholt 2013. OSCAR II LAND: Arnell & Mårtensson 1959 as Cephaloziella subdentata [cf. Frisvoll & Elvebakk 1996]. SABINE LAND: Crundwell 1978, Frisvoll & Elvebakk 1996. WEDEL JARLSBERG LAND: Frisvoll & Elvebakk 1996, Schuster & Konstantinova 1996.

*** Cephaloziella varians (Gottsche) Steph. View in CoL SVALBARD: Bryhn 1906 as Cephalozia divaricata var. incurva, Steere & Inoue 1978 as Cephaloziella arctica, Schuster 1980 as Cephaloziella arctica, Düll 1983 as Cephaloziella arctica, Söderström 1995 as Cephaloziella arctica, Damsholt 2002 , Söderström et al. 2002a, 2002b, Ellis et al. 2011, 2015a, Damsholt 2013, Hodgetts 2015, Mamontov & Vilnet 2017, Hodgetts & Lockhart 2020. BJØRNØYA: Berggren 1875a as Jungermannia divaricata var. incurva, Söderström 1995 as Cephaloziella arctica, Frisvoll & Elvebakk 1996 as Cephaloziella arctica, Damsholt 2002 , Söderström et al. 2002b. GUSTAV V LAND: TYPE of Jungermannia divaricata var. incurva, Berggren 1875a , 1875b both as Jungermannia divaricata var. incurva, Geissler & Bischler 1987 as Jungermannia divaricata var. incurva, Konstantinova & Savchenko 2011 , 2012. PRINS OSCARS LAND: Konstantinova & Savchenko 2008 a, Konstantinova et al. 2014, 2020b. BARENTSØYA: Philippi 1973 as Cephaloziella arctica, Konstantinova & Savchenko 2018 . EDGEØYA: Berggren 1875a as Jungermannia divaricata var. incurva, Philippi 1973 as Cephaloziella arctica, Heinemeijer 1979 as Cephaloziella arctica, Heinemeijer & van Dijk 2004 as Cephaloziella arctica . SPITSBERGEN: Persson 1942 as Cephaloziella alpina [Van Meijens Bay], Arnell & Mårtensson 1959 as Cephaloziella arctica [W. Spitsbergen]. ALBERT I LAND: SYNTYPE of Jungermannia divaricata var. incurva, Lindberg 1866 as Jungermannia divaricata var. incurva, Berggren 1875a as Jungermannia divaricata var. incurva, Geissler & Bischler 1987 as Jungermannia divaricata var. incurva, Frisvoll & Elvebakk 1996 as Cephaloziella arctica . DICKSON LAND: Konstantinova & Savchenko 2008 c, 2011. HAAKON VII LAND: Konstantinova & Savchenko 2006, Newsham & Goodall-Copestake 2021. HEER LAND: This publication. JAMES LAND: This publication. NATHORST LAND: This publication. NORDENSKIÖLD LAND: Eurola 1968 as Cephaloziella arctica, Hadač 1989 as Cephaloziella arctica, Konstantinova & Savchenko 2007 , 2008 a, 2008b, Konstantinova et al. 2020b. NY-FRIESLAND: SYNTYPE of Jungermannia divaricata var. incurva, Lindberg 1866 as Jungermannia divaricata var. incurva, Geissler & Bischler 1987 as Jungermannia divaricata var. incurva . OSCAR II LAND: Kobayashi et al. 1990 as Cephaloziella arctica, Newsham & Goodall-Copestake 2021 . PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. SABINE LAND: Philippi 1973 as Cephaloziella arctica . SØRKAPP LAND: This publication. TORELL LAND: Rejment-Grochowska 1967 as Cephaloziella arctica . WEDEL JARLSBERG LAND: Rejment-Grochowska 1967 as Cephaloziella arctica, Eurola 1968 as Cephaloziella arctica, Karczmarz & Święs 1988 , 1989, 1990a all as Cephaloziella arctica .

Clevea Lindb. View in CoL

*** Clevea hyalina (Sommerf.) Lindb. View in CoL [ Marchantia hyalina Sommerf. , Sauteria hyalina (Sommerf.) Lindb. View in CoL , Athalamia hyalina (Sommerf.) S.Hatt. View in CoL ]. SVALBARD: Berggren 1875 b, Kaalaas 1893a, Stephani 1898, Bryhn 1906, Herzog 1926, Arnell 1928, Jørgensen 1934, Müller 1952, Steere 1954, Arnell 1956, Worley 1972, Bonner 1977 as Clevea hyalina f. rufescens, Düll 1983 , Schuster 1992, Söderström 1995, Damsholt 2002, Söderström et al. 2002a, 2002 b, Long et al. 2003, Rubasinghe 2011, Damsholt 2013, Hodgetts 2015, Hodgetts & Lockhart 2020. BJØRNØYA: Sommerfeldt 1833, Berggren 1875a, Berggren 1875b, Frisvoll & Elvebakk 1996. GUSTAV V LAND: Berggren 1875 a, Konstantinova & Savchenko 2012 as Clevea hyalina var. rufescens . BARENTSØYA: Konstantinova & Savchenko 2018. SPITSBERGEN: Lindberg 1866 [Kingsbay]. DICKSON LAND: Frisvoll & Elvebakk 1996 as Athalamia hyalina, Belkina et al. 2015 View in CoL . HAAKON VII LAND: Arnell & Mårtensson 1959 as Clevea hyalina f. rufescens . JAMES LAND: Frisvoll 1981, Frisvoll & Elvebakk 1996. NORDENSKIÖLD LAND: This publication. NY-FRIESLAND: Berggren 1875a [rejected as Sauteria alpina View in CoL by Lindberg 1877]. OSCAR II LAND: Arnell & Mårtensson 1959 as Clevea hyalina f. rufescens . PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. SØRKAPP LAND: Dubiel & Olech 1990, Frisvoll & Elvebakk 1996. WEDEL JARLSBERG LAND: This publication.

Cryptocolea R.M.Schust. View in CoL

*** Cryptocolea imbricata R.M.Schust. View in CoL SVALBARD: Düll 1983, Schuster 1983, Davison 1993, Söderström 1995,

Checklist of liverworts on Svalbard 369

Schuster & Konstantinova 1996, Grolle & Long 2000, Konstantinova 2000, Damsholt 2002, Söderström et al. 2002a, 2002b, Damsholt 2013, Hodgetts 2015, Hodgetts & Lockhart 2020. ANDRÉE LAND: Frisvoll & Blom 1992, 1997, Konstantinova & Savchenko 2020b. DICKSON LAND: Frisvoll 1981, Frisvoll & Elvebakk 1996. OSCAR II LAND: Ellis et al. 2021. Diplophyllum (Dumort.) Dumort.

*** Diplophyllum albicans View in CoL (L.) Dumort. [ Jungermannia albicans L.]. SVALBARD: Berggren 1875b, Müller 1914b, Buch 1928, Frye & Clark 1946, Schuster & Damsholt 1974, Düll 1983, Miller et al. 1983 [with a ‘?’], Smith 1990, Söderström 1995, Bisang et al. 1998, Engel & Smith Merrill 1998, Paton 1999, Damsholt 2002, Söderström et al. 2002a, 2002 b, Konstantinova 2003, Vilnet et al. 2010, Heinrichs et al. 2012, Damsholt 2013, Hodgetts 2015, Bakalin & Vilnet 2018, Bakalin et al. 2020 a, Hodgetts & Lockhart 2020. BJØRNØYA: Summerhayes & Elton 1923a, Söderström 1995, Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997, Damsholt 2002, Söderström et al. 2002b. PRINS OSCARS LAND: Konstantinova & Savchenko 2008 a, 2015. ALBERT I LAND: Berggren 1875a, Schuster 1974, Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997, Konstantinova & Savchenko 2008a. NORDENSKIÖLD LAND: Konstantinova & Savchenko 2008a.

*** Diplophyllum taxifolium (Wahlenb.) Dumort. View in CoL [ Jungermannia taxifolia Wahlenb. ]. SVALBARD: Müller 1914b, Buch 1928, Jørgensen 1934, Frye & Clark 1946, Müller 1956a, Schuster 1974, Schuster & Damsholt 1974, Düll 1983 as Diplophyllum taxifolium var. macrostictum [with a ‘?’], Smith 1990, Söderström 1995, Frisvoll & Elvebakk 1996, Engel & Smith Merrill 1998, Paton 1999, Damsholt 2002, Söderström et al. 2002a, 2002b, Hodgetts 2015, Hodgetts & Lockhart 2020. PRINS OSCARS LAND: This publication. ALBERT I LAND: Berggren 1875a. HAAKON VII LAND: Arnell & Mårtensson 1959 as Diplophyllum taxifolium var. macrostictum . NOTE: Except for the new report here, Diplophyllum taxifolium View in CoL is only known from old sources.

Fuscocephaloziopsis Fulford View in CoL

*** Fuscocephaloziopsis albescens (Hook.) Váňa et L.Söderstr. View in CoL [ Pleuroclada albescens (Hook.) Spruce , Pleurocladula albescens (Hook.) Grolle ]. SVALBARD: Schofield & Crum 1972, Smith 1990, Söderström 1995, Paton 1999, Damsholt 2002, Söderström et al. 2002b, Vilnet et al. 2012a, Damsholt 2013, Feldberg et al. 2016. BJØRNØYA: Jørgensen 1934, Söderström 1995, Frisvoll & Elvebakk 1996, Damsholt 2002, Söderström et al. 2002b. PRINS OSCARS LAND: This publication. EDGEØYA: Heinemeijer 1979, Frisvoll & Elvebakk 1996. SPITSBERGEN: Philippi 1973 [Kvalvågen], Frisvoll & Elvebakk 1996 [Kvalvågen, Billefjorden]. ALBERT I LAND: Arnell & Mårtensson 1959 [“seems to be represent- ed mainly by its var. islandica ”], Frisvoll & Elvebakk 1996. DICKSON LAND: This publication. HAAKON VII LAND: Arnell & Mårtensson 1959 [‘seems to be represented mainly by its var. islandica ”]. NORDENSKIÖLD LAND: Arnell & Mårtensson 1959 [“seems to be represented mainly by its var. islandica ”], Frisvoll & Elvebakk 1996, Konstantinova & Savchenko 2008b, Prestø et al. 2014. OSCAR II LAND: Arnell & Mårtensson 1959 [‘seems to be represented mainly by its var. islandica ’]. PRINS KARLS FORLAND: Frisvoll & Elvebakk 1996 [s.lat.]. NOTE: All reports of Fuscocephaloziopsis albescens View in CoL should probably be referred to var. islandica View in CoL if the variety should be recognized at all. Frisvoll & Elvebakk (1996) rejected var. albescens completely from Svalbard.

* — var. albescens [ f. albescens ]. SVALBARD: Schuster 1974, Düll 1983 [with a ‘?’], Hodgetts 2015 [with a ‘?’].

* — var. islandica (Nees) Váňa et L.Söderstr. [ Jungermannia islandica Nees , Pleurocladula islandica (Nees) Grolle , Pleurocladula albescens var. islandica (Nees) L.Söderstr. et Váňa ]. SVALBARD: Berggren 1875b, Düll 1983, Söderström 1995, Söderström et al. 2002 a, Hodgetts 2015, Hodgetts & Lockhart 2020. BJØRNØYA: Berggren 1875a. ALBERT I LAND: Berggren 1875a. NORDENSKIÖLD LAND: Berggren 1875a. PRINS KARLS FORLAND: Berggren 1875a.

*** Fuscocephaloziopsis lunulifolia (Dumort.) Váňa et L.Söderstr. View in CoL [ Cephalozia lunulifolia (Dumort.) Dumort. ]. SVALBARD: Söderström 1995, Damsholt 2002, Söderström et al. 2002a, 2002 b, Konstantinova 2003, Damsholt 2013, Bakalin & Vilnet 2014, Hodgetts 2015, Hodgetts & Lockhart 2020. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967 as Cephalozia media, Karczmarz & Święs 1989 as Cephalozia media, Frisvoll & Blom 1992 , Frisvoll & Elvebakk 1996, Frisvoll & Blom 1997.

*** Fuscocephaloziopsis pleniceps (Austin) Váňa et L.Söderstr. var. pleniceps View in CoL [ Cephalozia pleniceps (Austin) Lindb. ]. SVALBARD: Smith 1990, Söderström 1995, Damsholt 2002, 2013, Söderström et al. 2002a, 2002b, Vilnet et al. 2012 a, Hodgetts 2015, Feldberg et al. 2016, Hodgetts & Lockhart 2020. PRINS OSCARS LAND: This publication. SPITSBERGEN: Persson 1942 [Kingsbay]. DICKSON LAND: Konstantinova & Savchenko 2008c. HAAKON VII LAND: Konstantinova & Savchenko 2006, 2008a. JAMES LAND: This publication, Frisvoll & Elvebakk 1996. NORDENSKIÖLD LAND: Hadač 1946, Arnell & Mårtensson 1959, Frisvoll & Elvebakk 1996, Konstantinova & Savchenko 2007, 2008 a, 2008b. OSCAR II LAND: Arnell & Mårtensson 1959, Kobayashi et al. 1990, Frisvoll & Elvebakk 1996. PRINS KARLS FORLAND: Konstantinova & Savchenko 2020a. SABINE LAND: Crundwell 1978, Frisvoll & Elvebakk 1996. WEDEL JARLSBERG LAND: Eurola 1968.

Gymnocolea (Dumort.) Dumort. View in CoL

*** Gymnocolea inflata (Huds.) Dumort. subsp. inflata View in CoL [ Jungermannia inflata Huds. ]. SVALBARD: Düll 1983, Geissler & Bischler 1987 as Jungermannia inflata var. rigidiuscula, Söderström 1995 , Paton 1999, Damsholt 2002, 2013, Söderström et al. 2002a, 2002b, Vilnet et al. 2010, 2012 a, Shaw et al. 2015, Hodgetts 2015, Mamontov & Vilnet 2017, Hodgetts & Lockhart 2020. PRINS OSCARS LAND: This publication, Konstantinova & Savchenko 2015. EDGEØYA: Heinemeijer 1979, Frisvoll & Elvebakk 1996, Heinemeijer & van Dijk 2004. SPITSBERGEN: Berggren 1875b. NORDENSKIÖLD LAND: TYPE of Jungermannia inflata var. rigidiuscula, Berggren 1875a as Jungermannia inflata var. rigidiuscula, Bonner 1976 as Jungermannia inflata var. rigidiuscula, Bednarek-Ochyra et al. 1987c , Hadač 1989, Frisvoll & Elvebakk 1996, Konstantinova & Savchenko 2008 a, 2008b, Konstantinova et al. 2020b. OSCAR II LAND: Inoue 1988, Kobayashi et al. 1990, Frisvoll & Elvebakk 1996, Newsham & Goodall-Copestake 2021. SABINE LAND: Philippi 1973, Frisvoll & Elvebakk 1996. WEDEL JARLSBERG LAND: Rejment-Grochowska 1967, Karczmarz & Święs 1989, Frisvoll & Elvebakk 1996.

Gymnomitrion Corda View in CoL

*** Gymnomitrion concinnatum (Lightf.) Corda View in CoL [ Cesius concinnatus (Lightf.) Gray View in CoL ‘ Cesia concinnata ’]. SVALBARD: Berggren 1875b, Müller 1909a, Herzog 1926, Arnell 1928,