Diastatea expansa, McVaugh, Bull. Torrey Bot.

2. DIASTATEA EXPANSA McVaugh, Bull. Torrey Bot. View in CoL Club 67: 787. 1940. TYPE: MÉXICO. Mexico: Correra , Dist. Temascaltepec, 1230 m, 19 Nov 1932, G. B. Hinton 2644 (holotype: GH!; isotypes: K-image!, NY-image!) .

Stems 7.5–47 cm tall, simple or with few to many ascending branches, 0.7–1.6 mm wide at base, winged; wings minutely ciliate. Leaves subsessile or pseudo-petiolate below, sessile above; blades green to purple, base truncate to cuneate and decurrent into narrow wings, apex acute; lowermost blades deltoid to elliptic, 2.3–12 mm long, 2.2–5.7 mm wide; middle to upper blades narrowly elliptic, 4.8–25 mm long, 1.2–5.2 mm wide; margins regularly serrate with prominent teeth; glabrous. Inflorescence strongly secund with few–20 flowers per stem, bracts narrowly elliptic to linear, 5.2–25 mm long, 0.8–3.1 mm wide, ascending to somewhat appressed. Flowers with pedicels 3–17 mm long in flower, glabrous, spreading to ascending, often bent distally in fruit; hypanthium oblique, obconic in flower, 0.6–1.5 mm in diameter, 0.5–0.7 mm in length from the calyx sinus to attachment point with pedicel, often expanded in fruit enveloping up to one-third of the mature capsule, glabrous; calyx lobes subequal, from one-half to three-quarters the length of the fruit, narrowly triangular to linear-elliptic, 1.2–3.3 mm long in flower, green or purple, margins entire or with 1–2 pair of teeth, glabrous; corolla purplish-blue, nectar guides yellow with raised projections at throat; tube funnelform, flaring from base to end of tube, and gibbous at base on ventral side, 3.0– 4.6 mm long from calyx sinus to lateral corolla sinus, 1.5–2.5 mm wide at distal end; dorsal lobes oblanceolate with a narrowed claw attaching distal portion to deltoid base, 2.7–4.0 mm long, 0.7–1.5 mm wide; ventral lobes ovate, obovate or oblong, with or without a mucronate tip, 3.2–4.7 mm long, 1.7–2.4 mm wide; filaments 3.0– 4.5 mm long; anthers 1.1–1.5 mm long, purple with downy white covering increasing at apex of dorsal anthers, plus two large, narrow trichomes and several slightly smaller secondary trichomes at apex of ventral anther pair. Fruits narrowly ellipsoid and oblique, 4.0–7.0 mm long, 2.0–3.0 mm wide, light brown seeds ellipsoid, 0.5 mm long, medium orange-brown. Figure 4B View FIG .

Chromosome Number —Unknown.

Distribution —Mexican states of Guerrero, Mexico, and Michoacan ( Fig. 7 View FIG ).

Notes — Diastatea expansa may be distinguished from D. tenera using both the leaf margins and the funnelform corolla as diagnostic traits. The corolla is not always as expanded as the specific epithet suggests. The leaf margins provide a reliable second diagnostic trait. Diastatea expansa has regularly serrate margins with very sharp and prominent teeth tipped with callosities while D. tenera has minute teeth that are often reduced to mere callosities.

Additional Specimens Examined — Mexico. — GUERRERO: Chirilagua , Dist. : Mina , 18 Nov 1936, Hinton 9852 ( MEXU). — MÉXICO: Moist ravine by Rıo Molino in pine-oak woods above reservoir, Valle del Bravo, 1–2 Nov 1949, Moore, Jr. & Cetto 5486 ( GH) ; Desviacion de la carretera Temazcaltepec-Tejupilco a Tenerıa y Rıo Grande, [18.9, -100.1], 16 Oct 1996, Vibrans 6271 ( MEXU) ; Zacazonapan, 3 Dec 1981, Rosiles M. s. n. ( MEXU, TEX). — MICHOACÁN: Zitacuaro-Zirahuato , Distr.: Zitacuaro, 21 Oct 1938, Hinton 13393 ( MEXU) ; 12 km al E de Uruapan , carretera a Patzcuaro, 23 Jan 1977, Rzedowski 34603 ( MEXU) ; Northwestern foothills of Cerro Tancıtaro, 13–14 km south of Periban de Ramos , 29 Nov 1970, McVaugh 24809 ( MEXU) ; A 3 km al SO de Ario de Rosales , carr. a La Huacana, 24 Oct 1981, Nu ~ nez 3437 ( MEXU) .