Spinarge miocenicus, Muller & André & Nel, 2025

Spinarge miocenicus n. sp.

( Figs 1; 2)

urn:lsid:zoobank.org:pub:

TYPE MATERIAL. — Holotype. France • body rather well-preserved with thorax, abdomen, one antenna, two complete forewings and partly preserved hind wings; Upper Miocene ; montagne d’Andance paleomaar, diatomites, Saint-Bauzile, Ardèche; MDA. R 2024.8.1.

TYPE STRATIGRAPHY AND LOCALITY. — Upper Miocene (7.54 Mya), montagne d’Andance paleomaar, diatomites, Saint-Bauzile, F-07210 Ardèche, France.

DIAGNOSIS. — In addition to the characters of the genus (see Discussion), third antennomere simple, elongate and slightly curved, 6.3× as long as wide; fore- and hind wings brown in basal half, more yellow in distal half.

DESCRIPTION

Head hidden in matrix; third antennomere elongate and slightly curved, 1.9 mm long, 0.3 mm wide. Thorax 3.4 mm long, 1.9 mm wide; mesoscutellar appendage fused to mesoscutellum. Forewing complete, 8.8 mm long, 3.1 mm wide; brown in basal half, more yellow in distal half; short anterior Sc branch (intercostal vein) located slightly anteriad Sc+R and 1M junction; pterostigma 1.5 mm long, 0.4 mm wide, strongly arched along cell 2R1, slightly arched along anterior wing margin; M+Cu slightly curved; 1M slightly longer than 1Cu, curved basally; R clearly deviating between junctions of Sc and 1M; fusion of R with M exceedingly short; R meeting 2M much less than halfway between short anterior branch of Sc and pterostigma; 2M slightly longer than 1rs-m; Rs emerging at pterostigmal base; 2Rs strongly curved, shorter than 3Rs; no r-rs crossvein; cell 2R1 with a distal abscissa ending in wing margin; cell 1R1 trapezoidal, 0.5 mm long; cell 1Rs 1.2 mm long, enclosed distally by subvertical 2rs-m (the latter slightly longer than 1rs-m); 3M shorter than 4M; cell 2Rs 1.5 mm long, distinctly wider along curved 3rs-m; 1Cu and 2Cu nearly straight, 1Cu as long as 2Cu; lm-cu meeting Cu at an angle of ca 90°, 1M and 1m-cu converging; cell 1Cu elongate, longer than cell R; 1cu-a (nervulus) located at middle of cell 1M, distinctly distad point of fusion of R with 1M; cell 2Cu long, trapezoidal; cell 2M fully enclosed distally by long and slightly sinusoidal 2m-cu; 1A long, nearly straight; basal anal cell 1A distally closed; a long fusion of 2A+3A with 1A; cell 2A quite long. Hind wings only partly visible, 6.8 mm long, brown in basal half and more yellow in distal half; cell R1 with a distal abscissa not ending in wing margin; cell Rs distinctly longer than cell 1M, both distally closed. Abdomen 5.7 mm long, 2.8 mm wide; fifth tergum with a long median process; genitalia not visible.