Empis (Polyblepharis) mongolica Shamshev

Empis (Polyblepharis) mongolica Shamshev , sp.n.

Figs 6–9.

TYPE MATERIAL. Holotype, ♂, labelled [in Cyrillic, Russian]: Mongolia, G.-Alt. aym. [=Gobi-Altayskiy aymak, now Govi-Altai aymag (province)], / Khasagt-Khayrkhan Ridge / 15 km S Dzhargalan / 14.viii.[1]970 Zaitzev ( ZISP, INS _DIP_0000628).

Paratypes. MONGOLIA. Govi-Altai aymag: same locality as holotype, 14.viii.1970 Nartshuk (1 ♂) . Bayankhongor aymag: southern slope of Ikh-Bogdo-Ul, N Bayan-Gobi, 2200– 2700 m, 7–8.viii.1969 Zaitzev (1 ♂, ZISP; dissected) .

DIAGNOSIS. Mid-sized species (body length about 6 mm); male eyes holoptic; palpus yellow; mesoscutum with 3 indistinct, brownish vittae; acrostichal and dorsocentral setae multiserial, pale in presutural part, laterotergite with pale setae; fore and mid legs mostly brownish, hind leg mostly brownish yellow; wing hyaline; abdominal tergites 2–5 in lateral view uniformly brownish pruinose; hypopygium large.

DESCRIPTION. Body length 6.0– 6.3 mm, wing 5.7–5.8 mm. Male ( Fig. 6). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny. Holoptic, upper ommatidia enlarged. Frons represented by very small subtriangular space just below ocellar triangle and larger space above antennae; bare. Face broad, bare (in one paratype 1 setula present just below antennae). Ocellar triangle with several moderately long, mostly pale (1–2 brownish to black setae usually present), fine setae; ocellars undifferentiated. Occiput mostly with numerous, pale, hair-like setae, bearing moderately long, strong, black setae dorsally; postoculars moderately long, fine, black and pale on upper part, very short laterally; postgena with pale hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than subglobular pedicel, both with very short setulae; postpedicel nearly 3X as long as basal width, with straight margins; stylus slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow, nearly 1.5X longer than eye height; palpus short, pale yellow, with rather numerous pale setulae.

Thorax black in ground-colour, mostly densely greyish pruinose; mesoscutum ( Fig. 7) with 3 indistinct, brownish vittae along acrostichal and dorsocentral setae. Antepronotum with 4–5 strong, black setae dorsally and long, hair-like setae laterally on each side (dorsal setae sometimes brownish yellow to pale). Postpronotal lobe with long, dense, fine, pale setae, 1 similar black seta. Proepisternum with tuft of numerous, pale, hair-like setae on lower part. Prosternum bare. Mesonotal setae: acrostichals pale and black on about anterior 1/3 of mesoscutum (sometimes mostly pale or mostly dark) and black posteriorly, arranged in 4–6 irregular rows, occupying anterior half of prescutellar depression, moderately long, dense; presutural dorsocentral setae black and pale, multiserial, similar to acrostichals; postsutural dorsocentrals black, becoming stronger, longer and less numerous towards scutellum, not separated by bare space from acrostichals on about anterior half of prescutellar depression, 3–4 prescutellars longest; presutural supra-alar space entirely covered with dense, long, pale, hair-like setae not separated by bare space from dorsocentrals (sometimes 1–2 black hair-like setae present); 4 black notopleurals (anterior seta usually slightly shorter); 1 similar postsutural supra-alar (with some additional short, fine, black and hair-like, pale setae); 1 long and 1 minute postalars, 4 scutellars of subequal lengths (lateral setae only slightly shorter than apicals). Laterotergite with numerous pale to pale yellow setae. Anterior and posterior spiracles pale yellow.

Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose; Leg colour: coxae and trochanters brown; fore and mid femora mostly brownish (yellowish at extreme apex and base; fore femur sometimes entirely brownish), hind femur brownish yellow; fore and mid tibiae mostly brownish (brownish yellow close to base), hind tibia brownish yellow (slightly darker at apex); fore and mid tarsi almost entirely brownish (only basitarsi paler at extreme base), hind basitarsus mostly brownish yellow (brownish at apex), remaining hind tarsomeres brownish. Fore coxa covered with pale, long, dense, hair-like setae anteriorly. All femora without whitish pilosity ventrally. Fore femur with row of mostly minute, fine anteroventral setae; covered with hair-like, moderately long, pale setae posteroventrally and posteriorly. Fore tibia with 3–5 short, anterodorsal setae (position and robustness variable). Mid femur with complete rows of equally short, strong anteroventral and posteroventral setae. Mid tibia slender; with 3–4 anterodorsal and 2 similar posterodorsal, moderately long setae. Hind femur slender; with row of 3–4 moderately long anterodorsal setae in about apical third; complete row of rather short and scattered anteroventral setae; fine short setae posteroventrally (longer close to base). Hind tibia simple, slender; with 7–9 moderately long, strong anterodorsal and 3–4 similar posterodorsal setae; simple setulae ventrally; 1 short seta in posteroapical comb. Tarsomeres of fore and mid legs slender, hind basitarsus slightly thickened at base; basitarsi with circlet of short, subapical setae, bearing scattered, spine-like setae ventrally; in addition, hind basitarsus with 2 short anterodorsal setae on about middle.

Wing membrane hyaline; entirely covered with microtrichia; veins mostly yellowish (costa between anterior margin of pterostigma and R 5 as well as apical portion of radial veins brownish to brownish yellow), well-sclerotized. Veins R 5 and M 1 divergent towards wing margin; radial fork acute; R 5 meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M 4 nearly 3X long- er than its middle portion. Pterostigma inconspicuous. Basal costal seta absent. Anal lobe well-developed; axillary incision very acute. Squama yellow, pale fringed. Halter yellow.

Abdomen entirely black in ground colour. Pruinescence: tergites 1–5 densely brownish grey pruinose, tergites 6–7 with yellowish brown iridescent spot dorsally, greyish laterally (dorsal view); sternites 1–7 densely greyish pruinose; sternite 8 faintly pruinose, contrastingly black (upper antero-lateral projection shiny). Chaetotaxy: tergites 1–5 covered with black, minute, numerous setulae dorsally, bearing pale, dense, hair-like and yellowish posteromarginal setae laterally; tergites 6–7 only with pale lateral setae; sternite 1 with 2 long, fine, pale setae, sternites 2–8 with numerous similar setae (slightly shorter on sternite 8), in addition, sternite 8 with moderately long, black posteromarginal setae. Pregenital segments: segment 6 unmodified; tergite 7 slightly concave posteriorly, sternite 7 with slightly produced postero-lateral corner, pleuron of segment 7 slightly sclerotised posteriorly; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 small, antero-lateral projections separated by depression (upper projection rounded, lower projection subconical) and large, broad projection on posterior margin of depression.

Hypopygium ( Figs 8, 9) large; epandrium and basal half of phallus brown, apical half of phallus and cerci brownish yellow; epandrial lamella shiny on middle part, with short pale and black setae along lower margin and near apex, bearing spinule-like setae at tip; cercus covered with dense setulae dorsally and near apex laterally. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella ( Fig. 8) subtriangular, with slightly produced and bent inward apex (lateral view). Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, subtriangular in ventral view; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus elongate ovate (lateral view), with apex extending to level of epandrial lamella apex, without dorsolateral, lobe-like projection in basal part (dorsal view), with serrate inner margin. Phallus ( Fig. 9) almost entirely hidden; zigzag bent, strongly constricted beyond middle; its basal portion gently curved, thick at base and becoming slightly narrower towards constriction; apical bend portion almost circular (its tip pointing to rear), long, almost uniformly slender (except slightly thickened extreme base). Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.

Female. Unknown.

DIFFERENTIAL DIAGNOSIS. The new species is similar to E. indigirca Chvála, 1999 (known from Eastern Siberia) and E. subcurta sp.n. described herein. Empis indigirca differs from E. mongolica sp.n. primarily by entirely black acrostichal and dorsocentral setae, faintly brownish infuscate wings, the presence of black posteromarginal setae on abdominal tergites 1–5 and by smaller male hypopygium [ Chvála, 1999]. Empis subcurta sp.n. differs from the new species primarily by entirely dark brown legs, entirely pale acrostichal and dorsocentral setae and smaller male hypopygium. In addition, in E. subcurta sp.n. strong mesonotal setae usually are yellow (except notopleurals) and abdominal tergites 1–5 are densely brownish pruinose dorsally and grey pruinose laterally (in dorsal view abdomen with broad brownish stripe).

ETYMOLOGY. The species is named after country of its origin ( Mongolia).

DISTRIBUTION. Mongolia.