Empis (Polyblepharis) tuvinica Shamshev

Empis (Polyblepharis) tuvinica Shamshev , sp.n.

Figs 29–31.

TYPE MATERIAL. Holotype, ♂ labelled: [ RUSSIA, Tuva] Turan [52°08′N 93°55E], Tuva / steppe slopes / V. Richter 1 VI [1]975 ( ZISP, INS _DIP_0000626). GoogleMaps

Paratypes. RUSSIA, Tuva: same data as holotype (1 ♂, 2 ♀♀, ZISP) GoogleMaps ; same locality as holotype, 5.vi.1975, V GoogleMaps . Richter (2 ♀♀, ZISP) ; same locality as holotype, 6.vi.1975, V GoogleMaps . Richter (1 ♀, ZISP) ; same locality as holotype, 7.vi.1975, V GoogleMaps . Richter (1 ♂, 2 ♀♀, ZISP) .

DIAGNOSIS. Mid-sized species (body length about 5 mm); male eyes dichoptic but frons very narrow at middle, narrow- er than anterior ocellus; postpedicel nearly 2.5X longer than basal width; labrum nearly 2X longer than eye height, palpus yellow; mesoscutum with 4 brown vittae, acrostichals biserial, dorsocentrals uniserial, laterotergite with mostly black setae; legs extensively yellow to brownish yellow; abdomen with sternite 8 yellow posteriorly, tergites 2–5 mostly shiny, tergites 6 and 7 with iridescent spots dorsally, tergites 1–5 bearing black posteromarginal setae laterally; female legs without pennate setae.

DESCRIPTION. Body length about 5 mm, wing 4.7–4.9 mm. Male ( Fig. 29). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; head setation mostly black. Eyes dichoptic, ommatidia equally small. Frons slightly broadened just below ocellar triangle, very narrow before middle (narrower than anterior ocellus), broadened towards antennae on remaining portion; bare. Face broad, bare. Ocellar triangle with 2 long and several short fine setae. Occiput with almost regular row of moderately long setae; postoculars moderately long on upper part and very short laterally; postgena with pale hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape slightly longer than subglobular pedicel, both with very short setulae; postpedicel nearly 2.5X as long as basal width, with straight margins; stylus slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow, nearly 2X longer than eye height; palpus short, mostly yellow, brownish close to base; with scattered, dark setulae.

Thorax black in ground-colour, mostly densely greyish pruinose and black setose; mesoscutum with 4 brown vittae (lateral vittae less distinct). Antepronotum with 4–5 black setae dorsally on each side. Postpronotal lobe with 1 long and several shorter setae of different lengths (some of them only slightly shorter than postpronotal seta). Proepisternum with tuft of several, pale, fine setae on lower part. Prosternum bare. Mesonotal setae: acrostichals arranged in 2 close irregular rows, short, absent on prescutellar depression; dorsocentral setae uniserial, presuturals rather long (slightly longer than antennal stylus), 3 long prescutellars; 1 presutural intra-alar (nearly as long as presutural dorsocentrals), 1 long presutural supra-alar, 3–4 notopleurals, 1 postsutural supra-alar (with 1–2 short, fine seta(e) anteriorly), 1 long and 1 minute postalars, 4 scutellars (lateral setae only slightly shorter than apicals); Laterotergite usually with a few strong black setae (sometimes some setae brownish to brownish yellow and additional short, fine, pale setulae present). Anterior and posterior spiracles yellow.

Legs with coxae densely greyish pruinose, remaining podomeres subshiny; mostly black setose; Leg colour: extensively yellow to brownish yellow; coxae (except apex), most part of fore femur (except apex and sometimes extreme base), tarsomere 1 at apex, most part of tarsomere 2 (except base) and entire tarsomeres 3–5 brownish. Fore coxa with scattered strong setae and pale setulae anteriorly. All femora whitish pilose ventrally. Fore femur with rows of minute anteroventral and posteroventral setae. Fore tibia with 2–4 short, anterodorsal setae (position and robustness variable). Mid femur with short setae in about apical half anterodorsally and anteriorly; bearing rows of minute, fine anteroventral and posteroventral setae becoming slightly longer and denser close to base; with similar setae on ventral face near base. Mid tibia rather slightly thickened; with 2–4 anterodorsal and 2–3 posterodorsal, short, strong setae (position and robustness variable). Hind femur slender; bearing complete row of short, numerous, anteroventral setae (longest setae shorter than half of femur maximal width); covered with short, finer setae posteroventrally. Hind tibia simple, slender; with 2–3 anterodorsal and 3 posterodorsal, moderately long setae; simple setulae ventrally; no seta in posteroapical comb. Tarsomeres of all legs slender; basitarsi with scattered, spine-like setae ventrally.

Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins yellowish brown, well-sclerotized. Veins R 5 and M 1 almost parallel towards wing margin; radial fork acute; R 5 meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M 4 nearly 3X longer than its middle portion. Pterostigma inconspicuous. Basal costal seta absent. Anal lobe well-developed; axillary incision nearly 90°. Squama yellow, pale fringed. Halter pale yellow.

Abdomen almost entirely brown in ground colour (tergites of pregenital segments somewhat darker), only sternite 8 yellow posteriorly (size of yellow space variable). Pruinescence: tergite 1 entirely densely greyish pruinose; tergites 2–5 mostly shiny (narrowly faintly pruinose anteriorly); tergites 6–7 with yellowish grey iridescent spot dorsally, shiny laterally (dorsal view); sternites 1–7 densely greyish pruinose, sternite 8 faintly pruinose. Chaetotaxy: tergites 1–5 with black and pale setulae dorsally and longer, fine, pale setae laterally, bearing long, black posteromarginal setae laterally (sometimes some posteromarginal setae brownish yellow to yellow); tergites 6–7 with scattered setulae; sternite 1 bare, sternites 2–7 with short, pale setae, sternite 8 with black setae (posteromarginals moderatelylong).Pregenitalsegments:segment 6unmodified;tergite 7 broadly concave posteriorly, sternite 7 unmodified; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subglobular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 small, antero-lateral projections separated by depression and larger, subconical projection on posterior margin of depression (slightly curved at apex).

Hypopygium ( Figs 30, 31) moderately large; yellow (epandrium mostly brownish yellow in holotype and one paratype); epandrial lamella with short, black setae more numerous along lower margin; cercus covered with black setulae. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella ( Fig. 30) subtriangular (lateral view). Hypandrium separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, subtriangular in ventral view; bare; gonocoxal apodeme small. Cerci separated with each other and from epandrium; cercus elongate ovate (lateral view), shorter than epandrial lamella, somewhat lobe-like produced at about basal half dorsally (dorsal view), with 2 small, pointed projections beyond middle of inner margin. Phallus ( Fig. 31) almost entirely hidden; zigzag bent, strongly constricted beyond middle; its basal portion gently curved, thick at base and becoming gradually narrower towards constriction; apical bend portion semicircular (its tip pointing to rear), rather mostly thicken, with short, slender subapical part. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.

Female. Similar to male except as follows. Frons broad, slightly broadened towards antennae; with scattered marginal setulae. Mid tibia slender; hind femur slightly thickened. Mid and hind femora mostly covered with minute setulae ventrally; hind femur with 4–5 short anteroventral setae in about apical 1/3. Abdomen almost entirely greyish pruinose; tergite 8 black, sternite 8 brownish yellow, both sclerites shiny. Cercus brown, short, covered with dense, pale, erect setulae.

DIFFERENTIAL DIAGNOSIS. The new species is similar to E. transbaicalica Shamshev and E. subtransbaicalica sp.n. described herein. Empis transbaicalica differs from E. tuvinica sp.n. primarily by broader frons of the male (slightly broader than anterior ocellus) and by the absence of an iridescent spot on the male abdominal tergite 6 [ Shamshev, 2006]. Empis subtransbaicalica sp.n. differs from E. tuvinica sp.n. primarily by pale laterotergal setae, 2 scutellar setae (sian 4) and pale setose abdomen.

ETYMOLOGY. The specific epithet refers to the territory of the origin of the new species, Tuva (Tyva) Republic of Russia Federation.

DISTRIBUTION. Russia (Tuva).