Empis (Polyblepharis) subtransbaicalica Shamshev

Empis (Polyblepharis) subtransbaicalica Shamshev , sp.n.

Figs 23–28.

TYPE MATERIAL. Holotype, ♂ labelled [in Cyrillic, Russian]: Mongolia, Ara-Khangayskiy / aymak [= Arkhangay aymag (province)], 30 km E / Tsetserleg [~ 47°28′N 101°27′E] / Nartshuk 2.4.[1]975 ( ZISP, INS _DIP_0000625). GoogleMaps

Paratypes. MONGOLIA, Töv aymag [= Tsentralny aymak (province)]: northern slope of Bogdo-ula near Ulan-Bator, 22.vi.1967, Zaitzev (3 ♂♂, 2 ♀♀) ; same locality, 29.vi.1967, Zaitzev (4 ♂♂, 2 ♀♀) ; same locality, 29.vi.1967, Kerzhner (1 ♀) ; northern slope of Bogdo-ula Mt., Zaysan , meadowsteppe slopes, 15.vi.1967, Kerzhner (3 ♂♂) ; same locality, 15.vi.1967, Zaitzev (5 ♂♂, 1 ♀) ; Ulan-Bator env., steppe, 24–29.vi.1970, Nartshuk (2 ♂♂, 1 ♀) ; same locality, 24–29. vi.1970, Kerzhner (2 ♀♀) ; same locality, 24–29.vi.1970, Kandybina (1 ♂, 2 ♀♀) . Uvs aymag [= Ubsunurskiy aymak (province)]: 5 km N of Khyargas, 10.viii.1970, Kerzhner (6 ♂♂, 3 ♀♀) ; same locality, 10.viii.1970, Nartshuk (1 ♀) ; same locality, 10.viii.1970, Kozlov (1 ♂, 1 ♀) ; Togtokhyn- Shil Ridge , 50 km ESE Ulangom, 7.viii.1970, Nartshuk (1 ♀) . Arkhangay aymag [=Ara-Khangayskiy aymak (province)] Tevshrulekh , 3.vii.1970, Kandybina (1 ♀) ; same locality, 24.vii.1970, Kandybina (1 ♀) . Govi-Altai aymag [=Gobi-Altayskiy aymak (province)]: Khasagt-Khayrkhan Ridge , 15 km S Dzhargalan, 14.viii.1970, Nartshuk (1 ♂) . RUSSIA. Buryatia: Okinskiy Stan, Verkhnie Sayany , 8.vii.1913, Tolstov (1 ♂) ; 10 km SW Selendum , pine forest, 24.vi.1971, V. Richter (1 ♂, dissected) . Irkutskaya Prov.: Baykal Lake, Chernorud , 27.vi.1966, A. Pleshanov (1 ♂) . Yakutia (Sakha): Mikhaylovka , 60 km NE Amga, 21.vi.1986, Bagachanova (1 ♂) ; same locality, 4.vi.1985, Maksimova (1 ♂, all paratypes in ZISP) .

DIAGNOSIS. Mid-sized species (body length 5–5.5 mm); male eyes dichoptic but frons very narrow at middle, narrower than anterior ocellus; postpedicel nearly 2.5X longer than basal width; labrum nearly 2.5X longer than eye height, palpus mostly yellow; mesoscutum with 4 brownish vittae, acrostichals biserial, dorsocentrals 1–2-serial, laterotergite with pale setae; legs extensively yellow, hind femur slender; abdomen mostly pale setose, tergites 2–5 faintly greyish pruinose dorsally; sternite 8 posteriorly and hypopygium yellow; female legs without pennate setae.

DESCRIPTION. Body length 5.1–5.6 mm, wing 6.1–6.5 mm. Male ( Fig. 23). Head capsule regions mostly densely greyish pruinose; face on lower margin and entire clypeus shiny; head setation mostly black. Eyes dichoptic, ommatidia equally small. Frons slightly broadened just below ocellar triangle, very narrow before middle (narrower than anterior ocellus), broadened towards antennae on remaining portion; bare. Face broad, bare. Ocellar triangle with 2 long and several short setae. Occiput with almost regular row of moderately long setae on each side; postoculars moderately long on upper part and short laterally; postgena with pale hair-like setae. Antenna with scape and pedicel brown, postpedicel and stylus black; scape nearly 1.5X longer than subglobular pedicel, both with very short setulae; postpedicel nearly 2.5X as long as basal width, with straight margins; stylus slightly longer than postpedicel basal width. Proboscis with labrum brownish yellow, nearly 2.5X longer than eye height; palpus short, mostly yellow, brownish close to base; with scattered, dark setulae.

Thorax black in ground-colour, mostly densely greyish pruinose and black setose; mesoscutum ( Fig. 24) with 4 brown vittae (lateral vittae less distinct). Antepronotum with 4–5 black to yellow setae dorsally on each side. Postpronotal lobe with 1 long and 3–4 shorter setae (sometimes with additional 3–4 yellow to pale setulae). Proepisternum with tuft of several pale, fine setae on lower part. Prosternum bare. Mesonotal setae: acrostichals arranged in 2 close irregular rows, short, absent on prescutellar depression; dorsocentrals uniserial, mostly short, 2 long prescutellars; 1 short presutural intra-alar, 1 long presutural supra-alar, 3–4 notopleurals (usually 3 strong, longer and 1 short, finer), 1 postsutural supra-alar (usually with 1 short, fine seta anteriorly), 1 long and 1 minute postalars, usually 2 scutellars (sometimes 1–2 additional, lateral, short, fine seta(e) present); sometimes 1–3 brownish yellow setulae in anterior part of notopleuron (usually absent). Laterotergite with a few long, strong, yellow and additional short, fine, pale setae. Anterior and posterior spiracles yellow.

Legs with coxae densely greyish pruinose, remaining podomeres subshiny; black setose; almost entirely yellow (including coxae), only tarsomere 1 at apex, most part of tarsomere 2 (except base) and entire tarsomere 3–5 brownish. Fore coxa with scattered setae anteriorly.All femora whitish pilose ventrally.Fore femur with rows of minute anteroventral and posteroventral setae. Fore tibia with 1–2 short, fine anterodorsal setae. Mid femur with row of mostly minute anteroventral setae (slightly longer and stronger near base) and more numerous, spinule-like setae posteroventrally (slightly longer in about basal half). Mid tibia with 2–3 anterodorsal and 3–4 posterodorsal, short setae (number, position and robustness variable). Hind femur slender; bearing complete row of rather short, numerous, anteroventral setae (longest setae nearly half as long as femur maximal width); covered with very short setae posteroventrally. Hind tibia simple, slender; with 3–4 anterodorsal and 3–4 posterodorsal, short setae; simple setulae ventrally; no seta in posteroapical comb.Tarsomeres of all legs slender; basitarsi with scattered, spine-like setae ventrally.

Wing membrane faintly brownish infuscate; entirely covered with microtrichia; veins yellowish brown, well-sclerotized. Veins R 5 and M 1 almost parallel towards wing margin; radial fork acute; R 5 meeting costa before wing apex; CuA+CuP complete (slightly weakened beyond middle). Cell dm moderately large, with strongly elongate apex; apical portion of M 4 nearly 3X longer than its middle portion. Pterostigma inconspicuous. Basal costal seta absent. Anal lobe well-developed; axillary incision nearly 90°. Squama yellow, pale fringed. Halter pale yellow.

Abdomen almost entirely brown in ground colour, only sternite 8 broadly yellow posteriorly (size of yellow space somewhat variable). Pruinescence: tergites 2–5 in lateral view mostly densely greyish pruinose (faintly pruinose along posterior margin), in dorsal view mostly faintly greyish pruinose; tergites 6–7 with yellowish grey iridescent spot dorsally, greyish pruinose laterally (dorsal view); tergite 1 and sternites 2–7 densely greyish pruinose, sternite 8 faintly pruinose. Chaetotaxy: tergites 1–7 mostly covered with numerous, very short, fine, pale setulae; tergites 1–3 with moderately long, yellowish, posteromarginal setae laterally (sometimes less distinct on tergite 3); tergite 8 bearing black setulae; sternite 1 bare, sternites 2–6 with pale setulae only, sternite 7 with additional moderately long, brownish to pale posteromarginal setae, sternite 8 covered with black setae only (posteromarginals moderately long). Pregenital segments: segment 6 unmodified; tergite 7 broadly concave posteriorly; sternite 7 with somewhat produced posterior corners; segment 8 with separated tergite and sternite; tergite 8 somewhat flattened, represented by two subtriangular sclerites separated mid-dorsally, with several short setae posteriorly; sternite 8 large, scoop-shaped, somewhat constricted anteriorly, with 2 antero-lateral projections separated by depression (upper projection slightly smaller and with bifid apex; lower projection larger, subconical) and small- er projection on posterior margin of depression.

Hypopygium ( Figs 25–28) moderately large; almost entirely yellow, only apical semicircular part of phallus yellow; epandrial lamella with short, black setae more numerous along lower margin; cercus covered with black setulae. Epandrium entirely broadly divided (epandrial bridge absent); epandrial lamella ( Fig. 25) rather subtriangular (lateral view), broadly rounded apically. Hypandrium ( Fig. 26) separated from epandrium; mostly membranous, undivided, narrowly sclerotized along margin, subtriangular in ventral view; bare; gonocoxal apodeme small. Cerci ( Fig. 27) separated with each other and from epandrium; cercus elongate ovate (lateral view), shorter than epandrial lamella, without additional lobe dorsally, serrate on about apical third of inner margin. Phallus ( Fig. 28) almost entirely hidden; zigzag bent, strongly constricted beyond middle; basal portion thick, gently curved; apical bend portion semicircular, thickened at base, with its tip pointing to rear. Ejaculatory apodeme moderately large, extended far beyond basal curvature of phallus, with lateral wings.

Female. Similar to male except as follows. Frons broad, parallel-sided, with scattered marginal setulae. Mid and hind femora mostly covered with simple setulae ventrally, hind femur only with 1–3 short, strong, anteroventral setae in about apical 1/3. Abdomen mostly densely greyish pruinose; mostly with pale setulae, yellow posteromarginal setae present only on tergites 1–2; tergite 7 faintly pruinose; tergite 8 yellow to brownish yellow, shiny; sternite 8 subshiny. Cercus brown, short, covered with dense, pale, erect setulae.

DIFFERENTIAL DIAGNOSIS. The new species is similar to E. transbaicalica Shamshev, 2006 known from the Eastern Siberia [ Shamshev, 2006]. Empis subtransbaicalica sp.n. differs from E. transbaicalica by pale laterotergal setae and pale setose abdomen. In addition, the male of the new species has pruinose abdominal tergites 2–5 (shiny in E. transbaicalica ) and spots of iridescent pruinosity on tergites 6 and 7 (only on tergite 7 in E. transbaicalica ).

ETYMOLOGY. The epithet of the new species refers to its similarity to E. transbaicalica .

DISTRIBUTION. Mongolia, Russia (Buryatia, Irkutskaya Province, Yakutia).