Prochremylus museumoceanus, Belokobylskij & Manukyan, 2025

Prochremylus museumoceanus Belokobylskij, sp. nov.

http://zoobank.org/ urn:lsid:zoobank.org:B0DBAF94-7433-4D84-ACE1-8D3B44E21E93

Figs 1–3 View FIGURE 1 View FIGURE 2 View FIGURE 3

Type material. Holotype: female (with syninclusions of Diptera ( Dolichopodidae and Ceratopogonidae ) and plantae—remains of Quercus spp. ), # 11367-37 (MWO).

Paratype: 1 female (with syninclusion: remains of Quercus spp. ), KAM # 78-24 (KAM).

Type locality and horizon. Baltic amber; Late Eocene, Prussian Formation, Priabonian range (37.2–33.9 Mya). Southeast coast of the Baltic Sea, Primorskiy quarry, Yantarnyi, Kaliningrad Province, Russia.

Etymology. This species name after Latin “museum” (= museum) and “oceanus” (= ocean) because the holotype of this species was found in the amber collection of Kaliningrad Museum of the World Ocean.

Description. Female (holotype) ( Figs 1 View FIGURE 1 , 2 View FIGURE 2 ). Body length 1.5 mm; fore wing length 1.2 mm.

Head (lateral view) 0.9 × as long as maximum height, 0.5 × as long as mesoscutum. Temple short, transverse diameter of eye (lateral view) ~ 3.5 × longer than temple. Eye 1.3 × as high as broad (lateral view). Malar space 0.2 × height of eye, almost equal to basal width of mandible. Maxillary palpus ~ 0.4 × as long as height of head.

Antenna weakly thickened, filiform, with rhinaria on antennomeres visible especially in apical half, with 12 antennomeres, weakly longer than head and mesosoma combined, 0.7 × as long as body. Scape long, ~ 2.0 × longer than maximum width, and 1.7 × longer than pedicel. First flagellomere ~ 4.0 × longer than its maximum width, almost as long as the widened second flagellomere; second flagellomere ~ 2.5 × longer than its maximum medial width. Penultimate flagellomere ~ 2.5 × longer than its maximum width, about as long as apical flagellomere.

Mesosoma relatively short, ~ 1.4 × longer than its maximum height. Median lobe of mesoscutum weakly protruding forward. Subalar depression shallow, relatively wide, rugose-reticulate. Precoxal sulcus distinct, relatively narrow, crenulate. Meso-metapleural suture distinct, crenulate. Propodeum with areas delineated by distinct carinae.

Wings. Fore wing wide, 2.6 × longer than its maximum width. Pterostigma 4.2 × longer than its width. Radial vein (r) arising before middle of pterostigma, inner distance of pterostigma from parastigma to radial vein (r) about 0.8 × inner distance from radial vein (r) to apex of pterostigma. First (r) and second (3RSa) radial abscissae forming distinct obtuse angle; first abscissa (r) 1.3 × longer than maximum width of pterostigma. Second radial abscissa (3- SR) 1.1 × longer than first abscissa (r), 0.3 × as long as the weakly curved third abscissa (SR1), almost as long as the straight first radiomedial vein (2-SR). Second radiomedial (submarginal) cell wide and short, weakly narrowed distally, ~ 2.0 × longer than its maximum width, 1.4 × longer than brachial (subdiscal) cell. Recurrent vein (m-cu) 0.6 × as long as first radiomedial vein (2-SR). Discoidal (discal) cell rather short and wide, ~ 1.5 × longer than its maximum width. Distance (1-CU1) between basal vein (1-M) and nervulus (cu-a) 0.8 × nervulus (cu-a) length. Brachial (second subdiscal) cell ~ 3.0 × longer than maximum width. Hind wing ~ 5.2 × longer than its maximum width. First abscissa of mediocubital vein (M+CU) 1.3 × longer than second abscissa (1-M).

Legs. Hind coxa ~ 1.7 × longer than wide. Hind femur (in weakly rotated fixation) about 4.0 × longer than its width. Hind tibia 1.3 × longer than hind femur, about equal to hind tarsus.

Metasoma 0.9 × as long as head and mesosoma combined; tergites behind first one rather soft, but second and third tergites relatively distinctly sclerotised. First metasomal tergite short, 0.7 × as long as second tergite, about as long as propodeum. Suture between second and third tergites distinct, relatively narrow and almost straight. Second tergite 1.7 × longer than third tergite. Ovipositor weakly widened subapically, then evenly taped towards apex. Ovipositor sheath 0.4 × as long as mesosoma, 0.8 × as long as hind femur, and 0.2 × as long as fore wing.

Sculpture and pubescence. Vertex and upper quarter of temple granulate-rugulose, lower part of temple smooth; face perhaps finely reticulate-punctate. Mesoscutum reticulate-granulate. Mesopleuron mainly smooth. Propodeum partly smooth basally and rugulose at least laterally and posteriorly, with areas delineated by coarse carinae; areola short and wide, perhaps smooth; basal carina rather long, about as long as areola. Hind coxa sculptured dorsally; hind femur smooth. First metasomal tergite mainly striate. Second and third tergites mainly smooth, with very sparsely but distinctly punctation; following tergites smooth. Hind tibia in dense and short setae. Ovipositor sheaths densely setose.

Colour. Body dark brown to partly black. Antenna and legs dark brown to black. Palpi brown. Fore wing faintly infuscate. Pterostigma entirely dark brown.

Paratype ( Fig. 3 View FIGURE 3 ). Differs from holotype by following characters.

Body length 1.3 mm; fore wing length 1.0 mm.

Head (dorsolateral view) ~ 1.5 × wider than maximum length. Transverse diameter of eye (sublateral view) ~ 4.2 × longer than temple. Ocelli small. Face ~ 0.7 × as wide as transverse diameter of eye. Maxillary palpus ~ 0.4 × as long as height of head.

Antenna with 11 antennomeres. Scape of antenna 2.3 × longer than its maximum width, 1.5 × longer than pedicel. First flagellomere ~ 3.6 × longer than its maximum width, 1.1 × longer than second flagellomere; second flagellomere ~ 2.8 × longer than its maximum medial width. Penultimate flagellomere ~ 3.0 × longer than its maximum width.

Mesosoma. Median lobe of mesoscutum not protruding forward. Precoxal sulcus oblique, situated in anterior half of lower part of mesopleuron. Propodeum with areas delineated by distinct carinae.

Wings. In fore wing, pterostigma 3.7 × longer than its width. Radial vein (r) arising weakly before middle of pterostigma, inner distance from parastigma to radial vein (r) about 0.9 × distance from radial vein (r) to apex of pterostigma. First abscissa of radial vein (r) weakly longer than maximum width of pterostigma. Second radial abscissa (3-SR) 1.3 × longer than first abscissa (r). Distance (1-CU1) between basal vein (1-M) and nervulus (cu-a) 0.7 × nervulus (cu-a) length. Brachial (second subdiscal) cell 3.3 × longer than maximum width. Hind wing 5.0 × longer than its maximum width.

Legs. Hind femur (in lateral fixation) 3.0 × longer than its width. Hind tibia 1.4 × longer than hind femur, equal to hind tarsus. Hind basitarsus 0.7 × as long as second to fifth segments combined.

Metasoma 0.8 × as long as head and mesosoma combined; tergites behind first one relatively soft. First metasomal tergite with high and widely separated dorsal carinae. Suture between second and third tergites narrow. Second tergite 2.0 × longer than third tergite. Ovipositor sheath 0.45 × as long as mesosoma, about as long as hind femur, 0.3 × as long as fore wing.

Sculpture and pubescence. Vertex finely striate laterally and perhaps reticulate medially, temple mainly smooth; frons smooth; face perhaps mainly smooth. Mesoscutum densely and distinctly reticulate-granulate. Propodeum partly smooth and rugulose at least laterally, with areas delineated by coarse carinae. Hind femur perhaps smooth. First metasomal tergite mainly striate. Second and third tergites smooth.

Colour. Body mainly black; metasoma medially faintly paler. Antenna and legs reddish brown to dark reddish brown, hind tibia mainly light brown, dark brown in distal 0.3. Palpi brown. Fore wing infuscate. Pterostigma entirely dark brown.

Male. Unknown.

Comparative diagnosis. This new species is similar to the type species Prochremylus brevicornis Brues, 1933 ; the differences between these taxa are shown in the key below.