Aulosaphes (Antiquasaphes) fumipennis Belokobylskij, 2025

Aulosaphes (Antiquasaphes) fumipennis Belokobylskij , sp. nov.

http://zoobank.org/ urn:lsid:zoobank.org:31747F5B-795F-45BE-92F5-A87C7D53EC14

Figs 4 View FIGURE 4 , 5 View FIGURE 5

Type material. Holotype: female (without syninclusions), KAM # 79-24 (KAM).

Type locality and horizon. Baltic amber; Late Eocene , Prussian Formation , Priabonian range (37.2–33.9 Mya). Southeast coast of the Baltic Sea, Primorskiy quarry, Yantarnyi, Kaliningrad Province, Russia.

Etymology. This species is named after Latin “fumis” (= smoke) and “pennis” (= pen, “wing”) referring to its infuscate fore wing.

Description. Female. Body length 2.2 mm; fore wing length 1.9 mm.

Head (dorsal view) 1.4 × longer than its medial length, about as wide as mesoscutum. Ocelli small. Temple long, curvedly narrowed behind eyes; transverse diameter of eye (lateral view) 1.1–1.2 × longer than temple in dorsal view, almost 3.0 × in lateral view. Eye 1.2 × as high as broad (lateral view). Face convex, its width 1.1 × larger than medial height. Malar suture distinct and complete. Malar space 0.2 × height of eye, almost equal to basal width of mandible. Hypoclypeal cavity subcircular, its width almost equal to distance from margin of cavity to margin of eye, 0.4 × width of face. Head below eyes (front view) distinctly and almost linearly narrowed. Maxillary palpus short, ~ 0.9 × as long as height of head.

Antenna slender, filiform, with 14 antennomeres, with numerous dense rhinaria on antennomeres especially visible in apical half, ~ 0.7 × as long as body. Scape rather long, ~ 1.6 × longer than maximum width, and 1.6 × longer than the rather narrow and long pedicel. First flagellomere ~ 4.5 × longer than its maximum width, almost as long as the widened second flagellomere; second flagellomere ~ 3.5 × longer than its maximum medial width. Penultimate flagellomere ~ 2.7 × longer than its maximum width, 0.9 × as long as apical flagellomere; later obtuse apically.

Mesosoma relatively short, ~ 1.5 × longer than its maximum height. Mesoscutum highly and subperpendicularly elevated above pronotum. Median lobe of mesoscutum weakly protruding forward. Notauli complete, deep, rather wide, crenulate. Prescutellar depression (scutellar sulcus) crenulate. Subalar depression rather deep, narrow, crenulate. Precoxal sulcus distinct, narrow and shallow anteriorly, deep and wide posteriorly, rugose-crenulate at least posteriorly. Meso-metapleural suture distinct, crenulate. Metapleural lower lobe long, rounded distally. Propodeum with areas delineated by distinct carinae, without postero-lateral tubercles.

Wings. Fore wing wide, 2.5 × longer than its maximum width. Pterostigma 3.4 × longer than its width. Radial vein (r) arising weakly before middle of pterostigma, inner distance of pterostigma from parastigma to radial vein (r) about 0.8 × its inner distance from radial vein (r) to apex of pterostigma. Radial (marginal) cell not shortened. Metacarp (1-R1) 1.2 × longer than pterostigma. First (r) and second (3-RS) radial abscissae forming subrectangular angle; first abscissa (r) 1.4 × longer than maximum width of pterostigma. Second radial abscissa (3-RS) 1.2 × longer than first abscissa (r), 0.5 × as long as the straight third abscissa (SR1), 0.9 × as long as the straight first radiomedial vein (2-SR). Second radiomedial (submarginal) cell wide and medium length, not narrowed distally, 2.5 × longer than its maximum width, 1.4 × longer than rather wide brachial (subdiscal) cell. Recurrent vein (m-cu) weakly postfurcal to first radiomedial vein (2-SR) and 0.6 × as long as this vein. Discoidal (discal) cell rather short and wide basally, ~ 1.5 × longer than its maximum width. Distance (1-CU1) between basal vein (1-M) and nervulus (cu-a) 0.8 × nervulus (cu-a) length. Mediocubital vein (M+CU1) weakly curved. Brachial (subdiscal) cell 3.5 × longer than maximum width. Parallel vein (CU1a) arising almost from middle of vein closed distal margin of brachial (subdiscal) cell (vein 3-CU1 almost equal to vein CU1b). Hind wing 3.7 × longer than its maximum width. Recurrent vein (m-cu) long, distinctly curved towards base of wing, sclerotised and brown.

Legs. Hind coxa ~ 1.4 × longer than wide. Hind femur slender, subclaviform, 5.0 × longer than its width in posterior quarter. Hind tibia rather distinctly widened, 1.1 × longer than hind tarsus. Hind basitarsus ~ 0.8 × as long as second to fifth segments combined; second segments of hind tarsus 0.3 × as long as basitarsus, ~ 1.1 × longer than fifth segments (without pretarsus).

Metasoma. First to third tergites combined 1.2 × longer than mesosoma. Rather soft segments behind third tergite distinctly protruding and strongly curved down. First metasomal tergite relatively short, strongly widened posteriorly, ~ 0.8 × as long as its posterior width, about as long as propodeum. Second tergite subsquare, about 1.1 × longer than its basal width, 1.4 × longer than third tergite. Suture between second and third tergites distinct, narrow, almost straight medially and weakly curved laterally. Third tergite weakly roundly curved posteriorly, not serrate in posterior margin, without distinct lamella or flange, without latero-ventral teeth. Ovipositor very weakly widened subapically, then evenly acuminate towards apex, 0.8 × as long as first to third tergites combined. Ovipositor sheath bad visible, perhaps shorter than second metasomal tergite.

Sculpture and pubescence. Vertex densely and distinctly transverse striate; frons rugose-striate; temple mainly smooth; face entirely distinctly reticulate-punctate. Mesoscutum entirely densely granulate. Scutellum perhaps weakly granulate. Mesopleuron mainly smooth. Metapleuron coarsely rugose-reticulate. Propodeum entirely rugose, with areas delineated by coarse carinae; antero-lateral areas relatively short and wide; areola short and wide; basal carina rather short, about 0.5 × as long as areola. Hind coxa rugose-striate at least dorsally and laterally; hind femur smooth, but with sparse punctation. First and second metasomal tergites distinctly and densely longitudinally striate. Third tergites densely and finely transversely aciculate, weakly granulate-reticulate antero-laterally; following tergites smooth. Hind tibia in dense and rather short setae.

Colour. Body dark brown to black. Antenna and legs black. Palpi brown. Fore wing distinctly and almost entirely infuscate. Pterostigma entirely dark brown; veins brown.

Male. Unknown.

Comparative diagnosis. This new species differs distinctly from all known extant species of the genus Aulosaphes ( van Achterberg 1995; Chen and He 1996; Belokobylskij 1998; He et al. 2000) by the characters listed in the subgenus diagnosis and in the key to subgenera.