Gregorites halogii Drohojowska & Szwedo, 2024

Gregorites halogii Drohojowska & Szwedo sp. nov.

Fig. 6 View Fig .

ZooBank LSID: urn:lsid:zoobank.org:act:BD32F78A-AD06-4C4D-A1CC-C8C536B50958 .

Etymology: Specific name after Hálogi, a jötunn (giant) and the personification of fire in Norse mythology.

Holotype: Male, NHMD 38960 [alternative ZMUC 902133 View Materials ], collected by C. V. Henningsen , 16.5.1956, deposited in Amber Collection in Natural History Museum of Denmark. Imago of whitefly is included in dark brown and small sized piece of amber (12×6× 2 mm) of flat cube shape. Type locality: Danish amber; western coast of Denmark.

Type horizon: Lutetian–Priabonian, Eocene.

Diagnosis.—Imago, male. Pedicel relatively short and thick with heart-shaped apical part and two short setae on lateral margin 4 th antennomere longer than 5 th, 6 th antennomere shorter than 7 th. Fore wing with vein CuA present. Apical part of claspers strongly curved dorsad, lateral margin with shallow indentation at base, medial margin with convexity in the middle part, claspers not crossed at apex. Aedeagus funnel-shaped at the base, further straight, with apex exceeding 3/4 of length of claspers.

Description. — Measurements: Table 1. Head from the dorsal side invisible ( Fig. 6A View Fig 7 View Fig ). The median incision in the anterior margin of head clearly visible from the ventral side ( Fig. 6A 6 View Fig , A 8 View Fig , A 9 View Fig , A 14). Compound eyes relatively large and bulging, not divided, ommatidia equal in size ( Fig. 6A View Fig 9 View Fig , A 14). Lateral ocelli invisible. Rostrum short and massive, reaching to the mesocoxa ( Fig. 6A 6 View Fig , A 8 View Fig , A 9 View Fig , A 14).

Antennae seven-segmented ( Fig. 6A View Fig 3 View Fig , A 9 View Fig , A 12, A 14). Scapus about 1.6 times as wide as long, lateral margins gently rounded. Pedicel relatively short (compared to the 2 nd antennomere of other described species) and massive, wider at the apical part, about 1.6 times as long as wide with anterior part heart-shaped and two short seta on lateral margin. Third antennomere the longest, about 1.5 times longer than 4 th. Fourth antennomere about 1.2 times longer than 5 th, 6 th antennomere the shortest, about 2.2 times shorter than 5 th, 7 th antennomere 1.5 times longer than 6 th, and tapering towards apex.

Thorax on the dorsal side completely covered by wings and an opaque layer of amber ( Fig. 6A View Fig 7 View Fig ). Fore wings visible only from the dorsal side. Fore wing gradually widening towards the apex, widest at about 3/4 of the length, almost 2 times as long as wide( Fig.6A View Fig 1 A View Fig 7 View Fig , A 13). Costal vein arcuate curved, anteroapical angle broadly rounded, apical margin and posteroapical angle rounded. Posterior margin of fore wing arched at the base, further almost straight. Common stem Sc+R+CuA short and thick. The Sc+R branch slightly arched, directed towards the anteroapical angle of the wing, forked before half its length. Branches R1 and Rs slightly arched, both not reaching a margin, Rs about 1.7 times as long as R1.

Vein CuA slightly wavy, protruding at an acute angle from the common stem Sc+R+CuA, not reaching the margin of wing. Claval vein CuP thin, slightly arched, reaching less than half the length of the wing, not reaching the margin. Hind wings invisible, covered by the fore wings Fig. 6A View Fig 7 View Fig ).

Prolegs the shortest, metalegs the longest, 1.2 times longer than mesolegs ( Fig. 6A View Fig 3 View Fig , A 4 View Fig , A 6 View Fig , A 8 View Fig ). Procoxae slightly conical, trochanters poorly marked, almost twice as wide as long, profemora shorter than tibia, protarsus two-segmented, basiprotarsomere visible, longer than apical protarsomere. Metatibia 2 times longer than metafemur. Basimetatarsomere about 1.6 times longer than apical metatarsomere. Total length of metatarsus similar to length of metafemur. Claws long, narrow and hooked, apical upwards ( Fig. 6A View Fig 3 View Fig , A 4 View Fig , A 6 View Fig , A 8 View Fig ). Paronychium invisible. Tibiae and tarsi covered with rows of setae, femora smooth. The comb of setae visible on the inner side of metatibia, consisting of long and evenly distributed setae ( Fig. 6A 6 View Fig ).

The abdomen invisible from the dorsal side, covered by wings and not transparent, cracked layer of amber ( Fig. 6A View Fig 7 View Fig ). From the ventral side abdomen seems quite massive, slightly longer than the total length of the head and thorax ( Fig. 6A 6 View Fig , A 8 View Fig , A 11 View Fig ). Three wax plates visible ( Fig. 6A View Fig 11 View Fig ). First wax plate partially visible, the same width as the 2 nd. Second and 3 rd wax plate as long as wide, with rounded edges. Pygofer concave in basal 2/3 of the length ( Fig. 6A View Fig 5 View Fig , A 10 View Fig , A 11 View Fig ). Aedeagus long (compared to the length of claspers), funnel-shaped at the base, further thin and straight, length of aedeagus reaches up to about 3/4 of the length of claspers ( Fig. 6A View Fig 5 View Fig , A 10 View Fig , A 11 View Fig ). Claspers elongated, broad, narrowed in the distal part, not crossed at apex, curved dorsad ( Fig. 6A View Fig 5 View Fig , A 10 View Fig , A 11 View Fig ). Lateral margin with slightly indentation at base, further arcuated. Medial margin with convexity in the middle part. Apical part of claspers covered with single setae.

Stratigraphic and geographic range.— Type locality and horizon only.

Gregorites ymiri Drohojowska & Szwedo sp. nov. Fig. 7 View Fig .

ZooBank LSID: urn:lsid:zoobank.org:act:F8035251-485E-4750-9D43-509D4CDAFC10 .

Etymology: After Ymir—in Norse mythology the first giant emerged from Ginnungagap.

Holotype: NHMD 38974 [alternative ZMUC 902147 View Materials ], collected by A.K. Andersen, 28.3.1968, deposited in Amber Collection in Natural History Museum of Denmark. Imago of whitefly is included in clear light brownish and moderate sized piece of amber (20× 10 mm) of irregular shape.

Type locality: Danish amber; western coast of Denmark.

Type horizon: Lutetian–Priabonian, Eocene.

Diagnosis.—Imago. Posterior margin of pronotum slightly arched, anterolateral angles acute, posterolateral angles rounded. Mesopraescutum wider than its length in the midline, anterior margin slightly curved, lateral angles rounded, the posterolateral margin gently curved. Anterior angles of mesoscutum strongly rounded, anterolateral margins with clear recesses, posterolateral margins slightly convex and curved, posterior margin with rounded recess. Anterior and posterior margins of the middle part of mesoscutellum mildly rounded, lateral parts almost straight. Anterior margin of mesopostnotum straight, lateral angles rounded, posterior angle curved. Metascutum with median indentation deep and curved, anterior margins rounded, posterolateral margins much longer than the anterior ones, rounded, side parts facing laterally, posterior margin with rounded recess. Anterior margin of metascutellum strongly rounded. Fore wing with vein CuA, branch Rs 2 times longer than branch R1.

Description. — Measurements: Table 1. Head partially visible ( Fig. 7A 7 View Fig , A 9 View Fig ). Compound eyes relatively large, anteriorly incised ( Fig. 7A View Fig 9 View Fig ). Ommatidia in the dorsal part smaller and densely arranged, in the ventral part are larger and sparsely arranged. Lateral ocelli not visible. Coronal suture not visible. The amber near the specimen is muddy, not transparent and not allowing to count the number of antennomeres ( Fig. 7A View Fig 4 View Fig , A 6 View Fig , A 9 View Fig ). Scapus is relatively large ( Fig. 7A View Fig 9 View Fig ), pedicel large and massive, widening towards apex. Third antennomere 2 times longer than 4 th. Other antennomeres not visible.

Thorax visible( Fig. 7A View Fig 3 View Fig , A 7 View Fig ). Pronotum wider than mesothorax and metathorax, seems to be also wider than the head. Lateral parts longer than length of pronotum in midline. Anterior margin slightly with acute protrusion in the middle, posterior margin slightly arcuate. Anterolateral angles acute, posterolateral angulately rounded. Triangular mesopraescutum clearly separated, wider than long in the midline. Anterior margin of mesopraescutum slightly arched, lateral angles rounded, posterolateral margins slightly arched, converging at an angle of 90°. M-shaped mesoscutum, narrower in the middle, with rounded anterior angles, about 6.8 times as wide as long in midline. Lateral parts form relatively long, narrow and blunt expansions. Anterolateral margins with clear indentations, posterolateral margins slightly convex and arched. Posterior margin concave in the middle. Mesoscutellum forms a narrow band, clearly wider and rounded in the middle part, the anterior and posterior margins of the mid part slightly rounded, lateral parts narrow and almost straight. Triangular mesopostnotum, slightly wider than its length in the midline, anterior margin straight, lateral angles rounded, posterolateral margins arcuate, posterior angle arched, triangular and prolonged caudad depression the midline visible. Metascutum partly covered by wings, strongly narrowed in the middle, several times as wide as long in the midline. Middle deep and arcuate indentation, anterior margins rounded, the lateral parts form narrow and blunt protuberances directed laterad, posterolateral margins longer than the anterior ones, rounded, posterior margin with a rounded indentation. Metascutellum narrow laterally, clearly wider in the middle, anterior margin rounded, posterior arcuate. Lateral bands narrow directed anteriad. The mesopostnotum, mesoscutellum and metascutellum measurements included only the middle part, without narrow lateral bands. Parapteron large and oval ( Fig. 7A View Fig 3 View Fig , A 7 View Fig ). Tegula relatively large, of irregular shape ( Fig. 7A View Fig 3 View Fig , A 7 View Fig ).

Fore wing widening apicad, the widest in about 2/3 of the length, about 2.3 times as long as wide ( Fig. 7A View Fig 1 View Fig , A 5 View Fig , A 8 View Fig ). Costal vein arcuate, anteroapical and posteroapical angles arcuate, apical margin arched. Posterior margin rounded basally, further straight in the middle part with slightly depression. Common stem Sc+R+CuA weakly visible. Branch Sc+R straight, directed apicad, forked about half the length of the wing. Branch R1 straight, not reaching the margin. Branch Rs almost straight, about 2 times longer than the branch R1, not reaching the margin. Vein CuA curved in the middle part, departing at an acute angle from the common stem Sc+R+CuA, not reaching the margin, almost parallel to the branch Rs. Claval vein CuP straight, not reaching the margin. Hind wing partially covered by the fore wings, however their shape can be described ( Fig. 7A View Fig 2 View Fig , A 5 View Fig , A 8 View Fig ). Hind wing widening apicad, the widest in about 3/5 of the length, about 2.6 time as long as wide. Basal part of the costal vein invisible, the anteroapical and posteroapical angles arcuate, the apical and posterior margins arched.

Legs visible only partly ( Fig. 7A View Fig 5 View Fig , A 9 View Fig ). Abdomen invisible. Gender cannot be determined ( Fig. 7A View Fig 5 View Fig ).

Remarks.—The specimen is placed right at the edge of amber, which had a significant impact on the poor quality of observations and photos taken.

Stratigraphic and geographic range.— Type locality and horizon only.