BALTOGNATHINA BRUNKE, ŻYŁA & SOLODOVNIKOV, 2019

SUBTRIBE † BALTOGNATHINA BRUNKE, ŻYŁA & SOLODOVNIKOV View in CoL SUBTRIB. NOV.

( FIGS 4–6)

urn:lsid:zoobank.org:act:7499F537-000C-4E9E-8295-E148E7D66EA8

Type genus

† Baltognathus Brunke, Żyła & Solodovnikov , here designated.

Diagnosis

† Baltognathina differs from all other subtribes of Staphylinini based on the following combination of character states: microsculpture of forebody entirely absent; labrum narrowly divided at middle; frontoclypeal puncture present; right mandible with single proximal tooth; dorsal basal ridge absent; pronotum without second puncture (sensu Schillhammer & Brunke, 2018) of dorsal row ( Fig. 4F); prosternum triangular, with lateral arms narrowed subapically ( Fig. 5C); scutellum with posterior scutellar ridge, and elytra with subbasal ridge extending horizontally to humerus and without any trace of a scutellar collar ( Fig. 5E, F), their epipleura without distinct row of coarse and impressed setose punctures ( Fig. 4D); mesotarsomeres setose dorsally; metacoxa without transverse carina.

Description

Head with non-geniculate antennae, inserted distant from anterior eye margin by a distance greater than twice the width of the antennal socket ( Fig. 4F); antennomeres 1–3 lacking dense tomentose pubescence and dense setation; apical antennomere compressed in narrow profile and without a dense field of microsetae; frontoclypeal puncture present; basal puncture present and not doubled; interocular punctures absent; parocular punctures absent; genal puncture absent; ventral basal ridge parallel with ventral part of post-occipital suture ( Fig. 5B); post-genal ridge present ( Fig. 5B); dorsal basal ridge absent; labrum not widely divided to base; second labial palpomere without dense brushes of setae; apical labial palpomere not greatly reduced in size, longer than previous segment ( Fig. 5A); gular sutures widely spaced and only weakly converging ( Fig. 5B); gula without distinct transverse basal impression; mandibles with curved outside edge, only a single tooth in proximal half.

Disc of head and pronotum entirely without microsculpture. Pronotum without second puncture (sensu Schillhammer & Brunke, 2018) in the discal row (only first and distal puncture present) ( Fig. 4F); flexible postcoxal process present, at base interrupted by inferior marginal line ( Fig. 5D); prosternum triangular, lateral arms narrowed at least subapically ( Fig. 5C); basisternum with pair of macrosetae present ( Fig. 5C); pronotum not fused to prosternum ( Fig. 5C). Mesoscutellum with posterior scutellar ridge present. Elytra with subbasal ridge extending horizontally to humerus ( Fig. 5E, F), the latter with row of humeral spines ( Fig. 5E); epipleuron with scattered setose punctures, without a distinct row of coarse impressed setose punctures ( Fig. 4D). Profemur with distinct apical row of lateroventral spines ( Fig. 6D); protarsomeres trapezoid, not cylindrical ( Fig. 6B), with tenent setae; meso- and metatarsi with five trapezoid segments ( Fig. 6C), setose dorsally; empodial setae present, subequally long on all tarsi; metacoxae without transverse carina. Hind-wing vein characters unknown. Protergal glands well developed ( Fig. 6A).

Abdominal tergites without accessory basal lines or posterior transverse basal lines; sternite III with basal transverse carina sharply produced posteriad at middle ( Fig. 6E).

Male primary and secondary sexual character states unknown.