Lamyctes Meinert, 1868

Lamyctes Meinert, 1868 View in CoL View at ENA

= Lamyctinus Silvestri, 1909

= Metalamyctes Verhoeff, 1941

Type species: Lamyctes fulvicornis Meinert, 1868 [= L. emarginatus ( Newport, 1844) ].

Assigned species: Lamyctes adisi Zalesskaja, 1994 ; Henicops africanus Porat, 1871 (= H. insignis Pocock, 1891 ); L. albipes Pocock, 1894 (=? L. mauriesi Demange, 1981 ); L. anderis Chamberlin, 1955 ; L. andinus Kraus, 1954 (= L. neglectus Chamberlin, 1955 ; L.

rectus Chamberlin, 1955;? L. subtropicalis Chamberlin, 1955 ); L. baeckstroemi Verhoeff, 1923 ; L. caducens Chamberlin, 1938 ; L. cairensis Chamberlin, 1921 ; L. calbucensis Ve rhoeff, 1939; L. castaneus Attems, 1909 ; L. cerronus Chamberlin, 1957 ; Lithobius coeculus Brölemann, 1889 ; Lamyctes cuzcotes Chamberlin, 1944 (= L. alancayanus Chamberlin, 1955 ); L. diffusus Chamberlin & Mulaik, 1940 ; Henicops emarginatus Newport, 1844 (= Lamyctes chathamensis Archey, 1917 ; L. fulvicornis Meinert, 1868 ; L. fulvicornis var. hawaiiensis Silvestri, 1904 ; L. kermadecensis Archey, 1917 ;? L. munianus Chamberlin, 1920 ; L. neozelanicus Archey, 1917 ;? L. navaianus Chamberlin, 1920 ; L. tasmanianus Chamberlin, 1920 ; L. zelandicus Chamberlin, 1920 ); L. gracilipes Takakuwa, 1940 ; L. guamus Chamberlin, 1946 ; L. guamus koshiyamai Shinohara, 1957 ; L. hellyeri n. sp.; Henicops inermipes Silvestri, 1897 ; L. insulanus Verhoeff, 1941 ; L. leon Chamberlin, 1944 ; L. leleupi Matic and Darabantu, 1977 ; L. liani Larwood, 1946 ; L. medius Chamberlin, 1951 ; L. microporus Attems, 1909 ; L. neglectus Lawrence, 1955 a (homonym of L. neglectus Chamberlin, 1955 ); L. neotropicus Turk, 1955 ; L. nesiotes Chamberlin, 1952 ; L. omissus Kraus, 1957 ; L. orthodox Chamberlin, 1951 ; L. oticus Archey, 1921 ; L. pachypes Takakuwa, 1941 ; L. pacificus Silvestri, 1905 [subspecies of L. inermipes (Silvestri, 1987) fide Demange and Silva, 1976]; L. pinampus Chamberlin, 1910 ; L. pius Chamberlin, 1911 ; L. remotior Chamberlin, 1955 ; L. robustus Lawrence, 1955 b; L. setigerus Lawrence, 1955 b; L. taulisensis Kraus, 1954 (= L. brattstroemi Chamberlin, 1955 ); L. tivius Chamberlin, 1911 ; L. tolucanus Chamberlin, 1943 ; L. transversus Chamberlin, 1962 ; Henicops tristani Pocock, 1893 .

Discussion: The phylogenetic analysis discussed below concludes that the absence of ocelli in Lamyctes hellyeri n. sp. has an independent origin from blindness in L. coeculus . Accordingly, these two species should not be united as a taxon defined by the absence of ocelli, as would be consistent with the Lamyctinus concept. Furthermore, the type species of Lamyctinus , L. coeculus , is nested within the cladistic structure of Lamyctes , rather than being its sister group. Recognising Lamyctinus as a separate genus (based on blindness and larval autapomorphies) renders Lamyctes paraphyletic. As such, we follow Brölemann (1930) and many subsequent workers in placing Lamyctinus in synonymy with Lamyctes .

Metalamyctes Verhoeff, 1941, defined by a bipartite first genital sternite in the male, may prove to be useful for a southern African/South Atlantic island clade that, in addition to the type ( Henicops africanus Porat, 1871 ) also includes Lamyctes baeckstroemi , L. castaneus , L. microporus , L. neglectus , L. robustus and L. tristani . Several other African species with strong projections on tergites 9, 11 and 13 were recognised by Chamberlin (1951) as forming the subgenera Lamyctes ( Eumyctes) and L. ( Neomyctes). Henicops sinuatus Porat, 1893 , the type species of Eumyctes, is more closely related to Lamyctopristus Attems, 1928 , than to Lamyctes (Edgecombe submitted), and we infer that other species referred to Eumyctes and Neomyctes by Chamberlin (1951) are likewise not members of Lamyctes . These species include the following: Lamyctes denticulatus Attems, 1907 ; L. ergus Chamberlin, 1951 ; and L. numidicus ( Latzel, 1886) .