Lamyctes Meinert, 1868

Lamyctes Meinert, 1868 View in CoL View at ENA

= Lamyctinus Silvestri, 1909

= Metalamyctes Verhoeff, 1941

Type species: Lamyctes fulvicornis Meinert, 1868 [= L. emarginatus ( Newport, 1844) ].

Assigned species: Lamyctes adisi Zalesskaja, 1994 ; Henicops africanus Porat, 1871 (= H. insignis Pocock, 1891 ); L. albipes Pocock, 1894 (=? L. mauriesi Demange, 1981 ); L. anderis Chamberlin, 1955 ; L. andinus Kraus, 1954 (= L. neglectus Chamberlin, 1955 ; L.

rectus Chamberlin, 1955;? L. subtropicalis Chamberlin, 1955 ); L. baeckstroemi Verhoeff, 1923 ; L. caducens Chamberlin, 1938 ; L. cairensis Chamberlin, 1921 ; L. calbucensis Ve rhoeff, 1939; L. castaneus Attems, 1909 ; L. cerronus Chamberlin, 1957 ; Lithobius coeculus Brölemann, 1889 ; Lamyctes cuzcotes Chamberlin, 1944 (= L. alancayanus Chamberlin, 1955 ); L. diffusus Chamberlin & Mulaik, 1940 ; Henicops emarginatus Newport, 1844 (= Lamyctes chathamensis Archey, 1917 ; L. fulvicornis Meinert, 1868 ; L. fulvicornis var. hawaiiensis Silvestri, 1904 ; L. kermadecensis Archey, 1917 ;? L. munianus Chamberlin, 1920 ; L. neozelanicus Archey, 1917 ;? L. navaianus Chamberlin, 1920 ; L. tasmanianus Chamberlin, 1920 ; L. zelandicus Chamberlin, 1920 ); L. gracilipes Takakuwa, 1940 ; L. guamus Chamberlin, 1946 ; L. guamus koshiyamai Shinohara, 1957 ; L. hellyeri n. sp.; Henicops inermipes Silvestri, 1897 ; L. insulanus Verhoeff, 1941 ; L. leon Chamberlin, 1944 ; L. leleupi Matic and Darabantu, 1977 ; L. liani Larwood, 1946 ; L. medius Chamberlin, 1951 ; L. microporus Attems, 1909 ; L. neglectus Lawrence, 1955 a (homonym of L. neglectus Chamberlin, 1955 ); L. neotropicus Turk, 1955 ; L. nesiotes Chamberlin, 1952 ; L. omissus Kraus, 1957 ; L. orthodox Chamberlin, 1951 ; L. oticus Archey, 1921 ; L. pachypes Takakuwa, 1941 ; L. pacificus Silvestri, 1905 [subspecies of L. inermipes (Silvestri, 1987) fide Demange and Silva, 1976]; L. pinampus Chamberlin, 1910 ; L. pius Chamberlin, 1911 ; L. remotior Chamberlin, 1955 ; L. robustus Lawrence, 1955 b; L. setigerus Lawrence, 1955 b; L. taulisensis Kraus, 1954 (= L. brattstroemi Chamberlin, 1955 ); L. tivius Chamberlin, 1911 ; L. tolucanus Chamberlin, 1943 ; L. transversus Chamberlin, 1962 ; Henicops tristani Pocock, 1893 .

Discussion: The phylogenetic analysis discussed below concludes that the absence of ocelli in Lamyctes hellyeri n. sp. has an independent origin from blindness in L. coeculus . Accordingly, these two species should not be united as a taxon defined by the absence of ocelli, as would be consistent with the Lamyctinus concept. Furthermore, the type species of Lamyctinus , L. coeculus , is nested within the cladistic structure of Lamyctes , rather than being its sister group. Recognising Lamyctinus as a separate genus (based on blindness and larval autapomorphies) renders Lamyctes paraphyletic. As such, we follow Brölemann (1930) and many subsequent workers in placing Lamyctinus in synonymy with Lamyctes .

Metalamyctes Verhoeff, 1941, defined by a bipartite first genital sternite in the male, may prove to be useful for a southern African/South Atlantic island clade that, in addition to the type ( Henicops africanus Porat, 1871 ) also includes Lamyctes baeckstroemi , L. castaneus , L. microporus , L. neglectus , L. robustus and L. tristani . Several other African species with strong projections on tergites 9, 11 and 13 were recognised by Chamberlin (1951) as forming the subgenera Lamyctes (Eumyctes) and L. (Neomyctes). Henicops sinuatus Porat, 1893 , the type species of Eumyctes, is more closely related to Lamyctopristus Attems, 1928 , than to Lamyctes (Edgecombe submitted), and we infer that other species referred to Eumyctes and Neomyctes by Chamberlin (1951) are likewise not members of Lamyctes . These species include the following: Lamyctes denticulatus Attems, 1907 ; L. ergus Chamberlin, 1951 ; and L. numidicus ( Latzel, 1886) .