Garnotia stricta var. longiseta, Hack.

c. var. longiseta Hack. View in CoL

Garnotia stricta Brongn.var. longiseta Hack.(1909a) 141; (1909b) no.744. — Garnotia mindanaensis Santos var. longiseta Santos (1950) 98. — Garnotia acutigluma (Steud.) Ohwi var. longiseta (Hack.) V.Prakash & S.K.Jain (1979) 6. — Type: Merrill in Kneucker Gram. Exsicc. 744 (holo W; B, BM, E, G, K, L, UC, US, WAG).

Garnotia stricta Brongn. var. adscendens Hack. (1909a) 141; (1909b) no. 744. — Lectotype: Hook.f. & Thomson s.n. (holo K; GOET,U), designated here (see note).

Muhlenbergia arundinella Ridl. (1913) 267 (‘ Muehlenbergia ’). — Lectotype: Boden Kloss s.n., Camp 6a (holo BM; K), designated here.

Garnotia ledermannii Pilg. (1914) 171. — Type: Ledermann 12521 (holo B lost).

Garnotia mezii Janowski (1921) View in CoL 86. — Lectotype: Schlechter 19542 (B lost, US fragm.; A, K, UC), designated by Gould (1972: 552).

Garnotia papuana Ohwi (1942) View in CoL 1. — Garnotia mezii Janowski var. papuana (Ohwi) Santos (1950) View in CoL 92. — Type: Kanehira & Hatusima 13092 (holo FU; A, BO, US fragm. in K).

Garnotia mindanaensis Santos (1943) View in CoL 135, t. 1. — Type: Bartlett 17235 (holo MICH; US).

Garnotia mezii Janowski var. maculata Santos (1950) 92. — Type: Carr 12395 (holo NY; BM, K 000943324 , KEP, L, US).

Garnotia mezii Janowski var. clemensiana Santos (1950) 93. — Type: Clemens 4878 (holo US; A, K).

Garnotia mezii Janowski var. longiramosa Santos (1950) 93. — Type: McGregor 74 (holo US).

Garnotia mindanaensis Santos forma volcanicola Santos (1950) 98. — Type: BS 75748 ( Ramos & Edaño) (holo NY; K, PNH lost).

Garnotia mindanaensis Santos var. ramosii Santos (1950) 99. — Type: BS 13983 ( Ramos ) (holo US; BM, K 0009433325 , PNH lost).

Garnotia stricta auct. non Brongn.

Plants perennial. Culms erect to erect and geniculate at base, simple to branched along the culm, 0.4–1 m long. Nodes glabrous, puberulous, or pilose. Sheath mouth glabrous to densely hairy. Ligule 0.2–0.5 mm long. Blades linear, flat to folded, 4–40 cm by 2–18 mm, glabrous to pilose with bulbous-based hairs, base truncate to narrowly attenuate. Panicle (10–) 20–40 cm long, common axis longer than the lowest branches, solitary to in whorls, at maturity appressed to spreading, scaberulous. Spikelets 2.5–5 mm long (excl. awn), callus hairs 0–0.5 mm long. First glume 2.5–5 mm long, shorter than to longer than the second glume, emarginate to mucronate. Second glume 2.5–5 mm long, apex acute to mucronate, arista 0–0.8 mm long. Lemma awn imperfect, column absent, arista thread-like, wavy, crinkly, or even looped, 6–15 mm long. Anthers 1–1.5 mm long.

Distribution — Malesia: N Sumatra ( Aceh, see note), Celebes (Minahassa, Makassar), Moluccas ( Ambon, Ceram, Halma- hera), Philippines ( Bohol, Leyte, Luzon, Mindanao, Visayan Isl.), New Guinea (Aru Isl.; widespread on the mainland; Bismarck Arch.), Australia (N Queensland).

Habitat — Primary forest, on rocks, abandoned fields, grasslands, road sides, shaded forest, etc. 0–3200 m altitude. It would seem to be a facultative rheophyte.

Collector’s notes — Growing in tussocks, culms leaning. Panicles narrow, more or less pendant, green, purplish green, purple. Anthers purple.

Notes — Garnotia ledermanii is only known from the type collection, apparently lost in Berlin, for no material has been seen by later authors. Santos (1950: 37) included it in his unranked Patulae Santos together with C. patula (Munro) Benth. We agree from the description with Gould (1972: 554) that it probably is a depauperate G. stricta var. longiseta .

On the same Kneucker sheet (and in the publication) the combination Garnotia stricta var. adscendens [Griseb.] Hack. was made based on G. adscendens [Munro] Griseb. (1868) 80, nom. nud. (based on Berghausia adscendens Munro (1865) 58 and G. stricta sensu Hook.f. (1896) 243: “quae differt arista inferne geniculata sub geniculo torta, foliis laterioribus”. Both names were not mentioned by Santos (1950), Bor (1960), or Chase & Niles (1962). Vouchers for Garnotia stricta var. breviaristata Stapf ex Hook.f. (incl. B. adscendens Munro var. breviaristata Munro and G. adscendens Munro var. breviaristata : Griffith 6783, Helfer s.n., Tenasserim, resp.) are in K and belong to G. tenella .

Setulose glumes occur occasionally, e.g. in Wisse 69 from the Minahassa (BO, WAG).