Bakyietaia liusanjieae Zhang, Yen & von Rintelen, 2025

Bakyietaia liusanjieae Zhang, Yen & von Rintelen gen. et sp. nov.

urn:lsid:zoobank.org:act:

Figs 24–25 View Fig View Fig

Diagnosis

Shell medium in size (25.69 ± 3.35 mm), sturdy, conical shell with lower spire, whorls not inflated; columellar lip recurved, forming callus covering umbilicus completely; operculum with small nuclear region (around 1/4 of inner opercular region) with shiny grains or veins; outer marginal tooth of radula with 26–27 small sharp cusps; distributed in northern Guangxi, China.

Etymology

Named after ‘Liu Sanjie’ (刘īÜ, Liu family’s third daughter), a legendary folk singer of the Zhuang people in Guangxi. The type locality of this species is close to her hometown. The recommended Chinese name is 歌üṉ田ª.

Material examined

Holotype

CHINA • spec. preserved in ethanol; Guangxi, Hechi City, Luocheng Mulao Autonomous County, near Hemu Town , river in Xinhui Village , 0.5 m deep on dam; 16 Mar. 2022; Le-Jia Zhang leg.; KIZ.240009 .

Paratypes

CHINA – Guangxi • 2 specs (preserved in ethanol); same data as for holotype; KIZ.240010 , KIZ.240011 • 2 specs (preserved in ethanol); same data as for holotype; ZMB.122722-2 • 2 specs (preserved dry); same data as for holotype; KIZ.240090 • 2 specs (preserved in ethanol); Guilin City, Baishou Town , Baishou River , 0.5 m deep on mud and sand substrate; 15 Mar. 2022; Le-Jia Zhang leg.; KIZ.240012 , KIZ.240013 • 2 specs (preserved in ethanol); same data as for preceding; ZMB.122738 • 1 spec.; same data as for preceding; KIZ.240091 • 1 spec. (preserved dry); Hechi City, Yizhou District , Liusanjie Town , on bank of Xiajian River ; 16 Mar. 2022; Le-Jia Zhang leg.; KIZ.240092 • 2 specs (preserved dry); same data as for preceding; 17 Mar. 2022; KIZ.240093 • 2 specs (preserved in ethanol); Hezhou City, Zhongshan County, Tonggu Town , Fengzhen Village , Shangpu , 0.5 m deep on mud substrate in a pond; 9 Mar. 2020; Shui-Yang Huang leg.; KIZ.240014 , KIZ.240015 • 1 spec. (preserved in ethanol); same data as for preceding; ZMB.122763 .

Other material

CHINA • 1 spec. (preserved dry); same data as for holotype; YHY-BAKLIU-XH-1.

Description

Adult shell ( Fig. 24A–K View Fig ) medium in size (25.69 ± 3.35 mm), thin but sturdy, conical shell with low spire, olive to olive brown, with five to six whorls, apex acute; teleoconch whorls not inflated, with three smooth relatively weak primary keels, with one to two even weaker smooth secondary keels located between suture and first keel, sometimes one additional weak keel located between first and second/ second and third keels, base of shell with one to five weak keels, all keels usually darker than rest of shell surface; aperture ovate, less than half of shell in height, inner shell surface bluish white, orange-red or dark purple, outer lip thin, always with a strong basal angular periphery, columellar lip covered with thick inner shell layers, white or orange red, adult columellar lip recurved, sometimes forming a relatively wide crescent-shaped callus, this structure or columellar lip completely covering umbilicus.

Operculum ( Fig. 24L–O View Fig ) corneous, ovate, relatively thick, dark orange to dark red; exterior surface smooth; interior surface with small nuclear region, around 1/4 of inner opercular region, with shiny grains or veins or smooth glossy surface.

Radular ( Fig. 25 View Fig ) (n = 3) central tooth with one broad central denticle and three to four small sharp cusps on either side; lateral tooth with one broad central denticle and two to three small sharp cusps on either side; inner marginal tooth narrow, about half of lateral tooth in width, with one broad central denticle and five small sharp cusps on either side; outer marginal tooth wide, almost same as lateral tooth in width, with 26 to 27 small sharp cusps.

Remarks

Bakyietaia liusanjieae gen. et sp. nov. can be distinguished from its congeners based on its medium-sized conical shell with whorls not inflated and lower spire. It differs from Bakyietaia liangzhuorum gen. et sp. nov. by having less prominent primary keels, thin outer lip and greater number of small cusps on outer marginal teeth (26–27 vs 20–25). The distributions of B. liusanjieae and B. liangzhuorum are allopatric (Northern Guangxi vs Southern Guangxi & West Guangdong).

Habitat and distribution

Rock substrate in the shallow water of rivers of northern Guangxi, China.