Lionepha sequoiae, (LINDROTH, 1963)

LIONEPHA SEQUOIAE ( LINDROTH, 1963) View in CoL

( FIGS 3B, 17C, D, 13C, D, 19B, 24B)

Bembidion sequoiae Lindroth, 1963: 260 View in CoL . Holotype ♂ in MCZ (type # 32532), examined. Type locality: Sequoia National Park , California.

Diagnosis: The most distinctive external feature of this moderately large Lionepha is the exceptionally small basal protarsomeres in males ( Fig. 19B), which are only slightly wider than the second protarsomeres, much smaller than in other Lionepha (see other images in Fig. 19). Elytra not iridescent, as sculpticells are not sufficiently transverse ( Fig. 17C, D). Aedeagus with broad apex, and with clearly visible dark scales on the internal sac membranes, yielding a speckled appearance ( Fig. 13C, D).

Additional characteristics: Body length 4.31–4.95 mm. Antenna piceous, with first antennomere rufous, at least on the underside. Legs rufous, with darker joints. Hind wings full-sized. Chromosomes of male 24 + X ( Table 5).

Distribution: This species has been found in the Sierra Nevada of California, the Cascades of Oregon and north to Vancouver, British Columbia ( Fig. 24B). There are, in addition, two low-elevation records from coastal California (Sonoma County); although one of these records (from 2.5 miles N of Cazadero on King Ridge, Big Austin Creek; CAS) is relatively recent, both records are unexpected and should be confirmed. Found from 30 to 2230 m in elevation. Specimens were collected from March through September.

Habitat: Found along the gravel and cobble shores of creeks in forests or in forest clearings. Over 80 specimens were found along partly shaded areas of School Creek in the Cascades of Oregon, with most found at the gravel and cobble edge of a splash pool below a culvert .