Coremacera catenata Loew, 1847

Coremacera catenata Loew, 1847 View in CoL

Figs 2 View Fig , 8 View Figs 6–9 , 17 View Figs 16–17

Material: Armenia, S of Lerik [38.76 ° N, 48.41 ° E], 24.06.1969, L. Zimina, 1♀ ( ZMUM) GoogleMaps ; Azerbaijan [on label as Armenia], Herher [39.71 ° N, 46.96 ° E], 13– 15.06.1956, L. Zimina, 4♀ ( ZMUM) GoogleMaps ; Georgia, Khashmi [41.75 ° N, 43.18 ° E], 31.07.1958, V. Zaitsev, 2♂, 1♀ ( ZIN) GoogleMaps ; Hungary, Bacs-Kiskun [county, ≈ 46.6 ° N, 19.4 ° E], poplar forest, 25.06.1970, K. Gorodkov, 2♂, 3♀ ( ZIN) GoogleMaps ; Iran: East Azerbaijan province, Tabriz [38.0 ° N, 46.3 ° E], 6.07.1914, Andrievsky, 1♀ ( ZIN) GoogleMaps ; Isfahan province, Ghomrood R., 33.42 ° N, 50.12 ° E, 21.05.2017, O. Kosterin, 1♂, 1♀ GoogleMaps ; Lorestan province, 16 km SE of Borujerd , 33.8 ° N, 48.9 ° E, 25.05.2017, O. Kosterin, 2♂, 1♀ GoogleMaps ; Markazi province: Arak env., 34.03 ° N, 49.75 ° E, 2000 m, 18– 30.05.2017, O. Kosterin, 4♂, 1♀ GoogleMaps ; 7 km NW of Shazand , 33.99 ° N, 49.36 ° E, 1850 m, 20.05.2017, O. Kosterin, 4♂, 1♀ GoogleMaps ; 9 km SW of Tafresh , 34.625 ° N, 49.947 ° E, 2300 m, 25.05.2017, O. Kosterin, 6♂, 2♀ (all ZMUM) GoogleMaps ; Moldova: Kishinev env., 1– 6.07.1970, R. Kamenskaya, 2♀ ; 26.06.1973, L. Zimina, 2♀; Vatichi [47.34 ° N, 28.61 ° E], 06–08.1953, 2♀ (all ZMUM) GoogleMaps ; Russia: Chechen Reg., Paraboch [43.47 ° N, 46.29 ° E], 13– 15.07.1927, Kirichenko, 2♂, 6♀ ( ZIN) GoogleMaps ; Crimea Reg.: Alma R. [≈ 44.84 ° N, 33.63 ° E], 10– 15.08.1899, Bashenov, 6♂, 4♀ ( ZIN) GoogleMaps ; Dagestan Reg.: Shura-Ozen River floodplain, 43.10 ° N, 47.46 ° E, 5.07.2015, D. Astakhov, 1♂ GoogleMaps ; North slope of Shalbuzdag Mt [≈ 41.4 ° N, 47.8 ° E], 22.07.1983, E. Narchuk, 1♂ (all ZIN) GoogleMaps ; Donetsk Reg., Volnovakha Distr. , 10 km E of Donskoe [47.50 ° N 37.65 ° E], 20– 31.08.2008, K. Tomkovich, 1♀ GoogleMaps ; Krasnodar Reg.: Dakhovskaya env., valley of Belaya R., 44.199 ° N, 40.170 ° E, 465 m, 17– 23.06.2009, K. Tomkovich, 1♀ ( ZMUM) GoogleMaps ; Lugansk Reg., Horodyshche [48.32 ° N, 38.64 ° E], 20.07.1952, B. Mamaev, 1♀ ( ZIN) GoogleMaps ; Samara Reg., Zhiguli [53.4 ° N, 49.3 ° E], Novoderzhkin, 3.07.1938, 1♂ GoogleMaps ; Chistovsky , 3.08.1950, 1♀ ( ZIN) ; Saratov Reg., Khvalynsky Nat. Park [52.48 ° N, 48.03 ° E], 27.06.2012, D. Astakhov, 1♂ ( ZIN) GoogleMaps ; Ulyanovsk Reg., Sengiley Hills Nat. Park , 54.0 ° N, 48.7 ° E, 16.07.2022, A. Nikolaeva, 1♂ ( ZMUM) GoogleMaps ; Volgograd Reg., Gorodishche [48.8 ° N, 44.5 ° E], steppe, 9.06.2012, D. Astakhov, 1♀ ( ZIN) GoogleMaps ; Kamyshin [50.1 ° N, 45.4 ° E], 25.07.1950, G. Viktorov, 1♂ ( ZMUM) GoogleMaps ; Zaporozhye Reg., Berdyansk env. [46.8 ° N, 36.8 ° E], 08.1954, 2♀ ( ZMUM) GoogleMaps ; Turkmenistan, Ahal Reg., Kopet-Dag, Koinekesir [38.22 ° N, 56.92 ° E], 25.06.1923, E. Smirnov, 1♀ ( ZMUM) GoogleMaps .

Distribution. Europe and South-West Asia. It is uncommon in Western Europe from where it has been recorded from the southern and central parts. It is more common in Eastern Europe, recorded from Moldova, Crimea, the Caucasus and the Volga region, along the Volga River it is recorded in the north up to 54 ° N. It is also not rare in South-West Asia: Turkey, Iraq, Iran, Israel, S-W Turkmenistan. It seems that C. catenata prefers rather arid regions.

Coremacera fabricii Rozkosny, 1981 Figs 3 View Fig , 7 View Figs 6–9 , 11, 12 View Figs 11–12

Coremacera cincta Fabricius, 1794 (preocc.)

Material: Russia: Moscow Reg.: Moskovsky env., Meshkovo , 55.59 ° N, 37.33 ° E, 11– 30.07.2017, K. Tomkovich, 3♂, 1♀ GoogleMaps ; Podolsk env., Vesennyaya , 55.39 ° N, 37.53 ° E, 29.05.2018, K. Tomkovich, 1♂ GoogleMaps ; Golitsino [55.6 ° N, 37.0 ° E] env., 7– 21.06.1980, A. Shatalkin, 2♂, 2♀ GoogleMaps ; Chashnikovo (56.04 ° N, 37.18 ° E) env., VI. 1973, (A. Shatalkin), 1♂ GoogleMaps ; Dmitrov distr., Kostino env. [56.316 ° N, 37.764 ° E], N. Vikhrev: 9– 10.06.2009, 2♂, 2♀ GoogleMaps ; 1.07.2009, 1♀ GoogleMaps ; 2.06.2010, 3♂ (all ZMUM) ; Smolensk Reg., Smolenskoye Poozerye National Park [55.5 ° N, 31.4 ° E]: 5– 8.06.1992, R. Zlobin, 2♂ GoogleMaps ; 13.06.1993, R. Zlobin ( ZIN) .

Distribution. Central Europe, rather uncommon species. To the north till Haapsalu, Estonia, 59 ° N ( Elberg 1968), to the east till Moscow region. In Central European Russia mostly collected in June.

Coremacera marginata marginata Fabricius, 1775

Figs 4 View Fig , 6 View Figs 6–9

Coremacera tristis Harris, 1780

Coremacera limbata Gmelin, 1790

Coremacera crinicornis Fallen, 1820 View in CoL Coremacera obscuripennis Loew, 1845 View in CoL syn. nov. Coremacera confluens Rondani, 1868 View in CoL syn. nov.

Material: Austria, Salzburg, Hallein [47.7 ° N, 13.1 ° E], 20– 24.07.2007, G. Pennards, 1♂, 1♀ ( ZMUM); Belarus: Gomel Reg., Mozyr env., 52.04 ° N, 29.32 ° E, 29– 31.07.2019, N. Vikhrev, 2♀; Vitebsk Reg.: Orsha env., 54.555 ° N 30.630 ° E, 28.07.2019, N. Vikhrev, 1♂ (all ZMUM); France: Provence-Alpes-Côte d'Azur region, Greolieres [43.795 ° N, 6.943 ° E], 24.08.1979, Heitmans, 1♂; Normandy, Seine Maritime foret de Brotonne [49.51 ° N, 0.79 ° E], 12.08.2009, G. Pennards, 1♀ ( ZMUM); Germany, Berlin, Wernsdorfer See [52.37 ° N, 13.66 ° E], K. Gorodkov, 21.06.1981, 3♀ ( ZIN). Greece, Kerkini, 41.21 ° N, 23.10 ° E, 1– 5.06.2007, G. Ramel, 1♀ ( ZMUM); Iran, Razavi Khorasan province, Mesched [36.3 ° N, 59.5 ° E], 15.09.1925, Jenikin, 1♀ ( ZIN, paratype of Coremacera marginata pontica Elberg, 1968 . This is the only known to me intermediate specimen, although it is more C. m. marginata than C. m. pontica.); Italy, Positano [40.63 ° N, 14.48 ° E], 2– 3.09.1925, El. Miram, 2♀ ( ZIN); Latvia, Tervete [56.48 ° N, 23.42 ° E], 7.07.1978, S. Kuznetsov, 1♂, 2♀ ( ZIN); Moldova: Bendery [46.8 ° N, 29.5 ° E], 27.07.1953, A. Zhelokhovtsev, 1♀; Kishinev env., 21.06.1970, R. Kamenskaya, 1♀ (both ZMUM); Kishinev [47 ° N, 29 ° E], 5.06.1981, V. Korenev, 1♂, 1♀ ( ZIN); Netherlands: Brabant, Oudland [51.55 ° N, 5.01 ° E], 2.06.2007, G. Pennards, 1♀; Meerssen [50.88 ° N, 5.75 ° E], 13.07.2009, G. Pennards, 1♂ ( ZMUM); Russia: Belgorod Reg., Les na Vorksle Nat. Res. [50.60 ° N, 35.98 ° E], 15.07.1976, Afanasieva, 1♂ ( ZIN); Kursk Reg.: Central Black Earth Nat. Res., oak forest [51.56 ° N, 36.08 ° E]: 24.06.1936, D. Dovnar, R. Zlobin, 4♂, 4♀ ( ZIN); Voronezh Reg., Liski env., Divnogorye [50.97 ° N, 39.32 ° E], 4– 22.07.1994, R. Zlobin, 2♂, 1♀ ( ZIN); Serbia, Crni Vrh, 43.407 ° N, 22.587 ° E, 800 m, 1– 7.07.2015, N. Vikhrev, 1♂, 1♀; A. Ozerov and M. Krivosheina, 1♂, 1♀; UK, Hartslock [51.5 ° N, 1.12 ° W], 6– 7.07.1999, C. Raper, 3♂; Ukraine: Kropivnytsky Reg., Elizavetgrad [Kropivnytsky, 48.5 ° N, 32.3 ° E], 5.07, E. Yatsentkovsky, 1♀ ( ZIN); Zakarpatsky Reg.: 15 km S of Rakhiv [47.92 ° N, 24.17 ° E], 14.07.1964, L. Zimina, 2♀; 25 km N of Rakhiv [48.26 ° N, 24.35 ° E], 5.08.1964, L. Zimina, 2♀ (both ZMUM); Kiev Reg., Kiev, 10.08.1902, Yu. Vagner, 1♀; Odessa Reg., Odessa [46.5 ° N, 30.7 ° E], 7.08.1978, Yu. Verves, 1♂; Poltava Reg., Hadiach [50.37 ° N, 33.98 ° E], Knipovich, 1♀ (all ZIN).

Distribution. W Palaearctic. From ~ 60 ° N to ~ 40 ° N in W Europe and 46 ° N in E Europe. I have only seen in ZMUM and ZIN specimens collected west of the Volga River. Elberg (1968) listed specimens collected in Kazan (i.e. on the eastern bank of Volga) and from the western Urals, Perm region, Kungur, K. Borisova [Uchleskhoz, Preduralie NP, presently abolished, 57.36 ° N, 57.14 ° E]. However, these specimens were not found in ZIN where they should be stored. Anyway, C. m. marginata is unknown in Siberia, but appears to be present in South-West Asia: Turkey, Bolu [40.7 ° N, 31.6 ° E] and Hakkari [37.5 ° N, 43.7 ° E] (as C. obscuripennis ) ( Leclercq & Schacht 1986) and Iran, Mesched [36.3 ° N, 59.5 ° E] (as C. m. pontica) ( Elberg 1968; Vikhrev et al. 2023). Recently

Khaghaninia et al. (2018) reported C. marginata from East Azerbaijan province of Iran, unfortunately without indication which subspecies was found.

Coremacera marginata pontica Elberg, 1968

Material: Armenia, Tsaghkadzor [40.53 ° N, 44.73 ° E, 1900 m], 15.07.1955, L. Zimina, 1♂; Azerbaijan, Nakhchivan Reg. , Orduban [38.9 ° N, 46.0 ° E, 800 m], 31.07.1970, V. Rikhter, 1♂ ( ZIN) GoogleMaps ; Georgia, Borjomi [41.84 ° N, 43.39 ° E, 900 m], 1867, Brandt, 2♂, 1♀ ( ZIN) GoogleMaps ; Russia: Chechen Reg., Vedeno [42.96 ° N, 46.10 ° E, 750 m], 5.06.1972, V. Rikhter, 2♂, 1♀ ( ZIN) GoogleMaps ; Crimea Reg.: Alushta env., Crimean Nature Reserve [≈ 44.7 ° N, 34.3 ° E], 9.08.1955, B. Rodendorf, 1♂, 1♀; 12.07.1956, B. Rodendorf, 1♂, 3♀; Pionerskoe [44.88 ° N, 34.19 ° E]: 4– 9.07.1976, L. Zimina, 2♂, 1♀; 15.06.1975, A. Zhelokhovtsev, 1♂; Kara-Dag [44.93 ° N, 35.23 ° E], 22.06.1982, L. Zimina, 1♀; Morskoe [44.83 ° N, 34.82 ° E], 13.08.2007, K. Tomkovich, 1♂ (all ZMUM) GoogleMaps ; Alma R. [≈ 44.84 ° N, 33.63 ° E], 10.08.1899, Bashenov, 5♂, 1♀ ( ZIN) GoogleMaps ; Dagestan Reg. , 15 km SW of Sergokala [≈ 42.33 ° N, 47.55 ° E], 15.07.1983, E. Narchuk, 15♂, 7♀ ( ZIN) GoogleMaps ; Kabardino-Balkaria Reg., Nalchik [43.5 ° N, 43.7 ° E], 12.07.1915, Golovleva & Kirichenko, 1♀ ( ZIN) GoogleMaps ; Krasnodar Reg.: Goryachy Klyuch [44.65 ° N, 39.15 ° E], 11.06.1997, K. Tomkovich, 1♀; Novorossiysk env., S Ozereevka [44.68 ° N, 37.63 ° E], 16.06.2001, K. Tomkovich, 1♂; Dakhovskaya env., valley of Belaya R., 44.199 ° N, 40.170 ° E, 465 m, 17– 23.06.2009, K. Tomkovich, 1♀; 18– 31.08.2009, K. Tomkovich, 2♂; ( ZMUM) GoogleMaps ; Osetia-Alania Reg.: Sukhotskoe [43.68 ° N, 44.44 ° E], 2.08.1988, A. Ozerov, 1♀; Tsey [42.79 ° N, 43.91 ° E, 1800 m], 16.09.1989, A. Shatalkin, 1♂ GoogleMaps ; Stavropol Reg., Shpakovskoye [= Mikhaylovsk, 45.1 ° N, 42.0 ° E], 13.06.1988, S. Belokobylsky, 1♀ ( ZIN) GoogleMaps ; Essentuki env. [44.0 ° N, 42.9 ° E], no date, E. Shirokova, 2♂ ZMUM GoogleMaps ); Volgograd reg., Sarepta [48.52 ° N, 44.51 ° E], T. Bekker, 1868, 1♂, 2♀ ( ZIN) GoogleMaps .

Distribution. Crimea and Caucasus, the latter also includes the lowlands adjacent to the north, such as the Stavropol and Volgograd regions. It is noteworthy that for the Black Sea territories west of Crimea (i.e. Odessa region and Moldova) only C. m. marginata was collected. Similar situation is in the Transcaucasian region: Armenia, Azerbaijan, Georgia, Dagestan — only C. m. pontica while in Turkey — only C. m. marginata . Thus, as far as we presently know C. m. pontica and C. m. marginata are nowhere sympatric, as it should be for subspecies of the same species by definition.

Coremacera turkestanica Elberg, 1968 is transferred to another genus, see Euthycera turkestanica in Chapter II.

Coremacera ussuriensis Elberg, 1968 View in CoL Figs 5, 9

Type locality: Russia, Primorsky region, Lebehe [presently Tikhovodnoe, 44.44 ° N, 132.57 ° E] GoogleMaps .

Material: Russia: Jewish Reg., Listvyanka R. (≈ 48.10 ° N, 132.76 ° E) GoogleMaps , 12.08.2003, I. Melnik, 1♀ ( ZMUM) ; Primorsky Reg., 20 km S of Spassk-Dalny, Evseevka [44.4 ° N, 132.9 ° E] GoogleMaps , 9.07.1993, S. Belokobylsky, 1♂ ( ZIN); 30 km NW of Arseniev [44.29 ° N, 132.95 ° E], on light GoogleMaps , 31.08.1999, Mironov, 1♀ ( ZIN); Kamenushka vill. [43.62 ° N, 132.23 ° E]: 14.07 GoogleMaps – 1.08.1983, A. Shatalkin, 5♂, 3♀; 6– 20.08.1983, A. Ozerov, 2♂, 2♀; 15.07– 13.08.1984, A. Shatalkin, 2♂, 4♀; 13– 15.08.1987, A. Shatalkin, 3♀; 19– 23.08.1989, S. Churkin, 2♂, 1♀ ( ZMUM); Kedrovaya Pad’ Nat. Res. , [43.1 ° N, 131.4 ° E] GoogleMaps : 23.07.1984, A. Shatalkin, 2♀ ( ZMUM) .

Distribution. So far known only from the Russian Far East.

Remarks. The holotype was not found in ZIN collection where it should be stored. Presumably it is still in Tartu, Estonia, where Elberg worked.