Thyanta (Argosoma) boliviensis, Rider, 1991

Thyanta (Argosoma) boliviensis Rider, new species

Figs. 230-244, Map 3

Description. Medium to large; dorsal surface olive green to reddish-brown; often with reddish-purple markings between humeral angles, on dorsal surface of head, and on apex of scutellum; punctures concolorous with surface.

Apex of head evenly rounded; outer jugal margins sinuous ( Fig. 231). Punctures on head rather dense, surface sometimes appearing reticulate. Antennae green to pale brown, distal 3 segments usually marked with red. Anterolateral margins ofpronotum straight in dorsal view; humeral angles rounded to nearly angulate, produced beyond base of adjacent coria by one-half width of eye or less ( Fig. 230). Pronotal cicatrices immaculate. Hemelytral punctures shallow, slightly more dense on exocorium; posterior margins slightly convex; costal angles narrowly rounded, usually reaching to middle of penultimate connexival segments ( Fig. 230); hemelytral membranes hyaline, usually lacking all brown flecks. Connexiva narrowly exposed, posterolateral angles of segments piceous.

Ventral surface yellowish-green to brown; punctures concolorous with surface. Rostrum pale brown, apical half of segment 4 piceous, reaching onto base of third (second visible) abdominal segment. Ostiolar canals acuminate apically. Femora and tibiae pale brown, tarsal segments and distal third of each tibia sometimes darker. Postspiracular spots lacking; posterolateral angles of abdominal stemites piceous.

Mesial margins of basal plates convex, separated basally and distally; posterior margins straight to slightly convex; posteromesial angles slightly emarginate ( Fig. 242). Sclerotized rod elongate, neither swollen subapically nor abruptly narrowed apically; dilation of spermatheca extending about three-fourths length of sclerotized rod, not abruptly narrowed on apical fourth ( Fig. 243); spermathecal bulb slightly elongate, spermathecal duct with small amount of coiling below proximal flange ( Fig. 244). Posterior margin of pygophore sinuously U-shaped in caudal view, medial portion slightly concave ( Fig. 238); pygophore emarginate in lateral view ( Fig. 241); posterolateral angles not distinctly prominent in ventral or dorsal views ( Figs. 239. 240). Apex ofeach paramere acute, nearly spinose in medial view ( Fig. 232); paramere slightly lunate in ectal view, apex nearly spinose ( Fig. 234); roughened, spiculate area on lateral surface ofparamere linear in shape ( Fig. 223), corresponding black carina on wall ofpygophore also linear. Each lateral conjunctival lobe ofaedeagus with one acute diverticulum apically and one obtuse slightly sclerotized diverticulum ventrally ( Fig. 235); dorsomedial lobe well developed ( Fig. 236); penisfilum and median penial lobes nearly hidden by conjunctiva ( Fig. 237).

Measurements. Total length 7.41-9.90 (7.73); total width 4.73-6.07 (4.89); medial length of pronotum 1.40- 1.82 (1.51). Medial length of scutellum 3.05-4.08 (3.13); basal width 3.02-3.86 (3.13); width at distal end offrena 0.99-1.32 (1.03). Length ofhead 1.57- 1.82 (l.64); width 2.03-2.32 (2.12). Length of segments 1 -5 of antennae 0.48-0.55 (0.52), 0.75-0.99 (0.81), 1.10-1.32 (1.25), 1.32-1.53 (1.47), and 1.36-1.44 (1.44), respectively. Length of segments 2-4 ofrostrum 1.21-1.51 (1.21), 0.81-0.96 (0.85), and 0.81-0.99 (0.92), respectively.

Holotype. <3 labeled (a) " Yungas de La Paz , Bolivia Dec. 4-20, 1955, 1200-1700 M Luis E. Pena, Collector" (b) "JC Lutz Collection 1961." Deposited in the U.S. National Museum of Natural History (Washington, D.C.) .

Paratypes. 4455, 6622. Labeled same as holotype (455 222 USNM) ; labeled as holotype, except lacking (b) (655 422 AMNH, FSCA) ; labeled as holotype, except (b) " Thyanta hurneralis Ruckes Det. J. C. Lutz" (5 AMNH) ; labeled as holotype, except (b) " Thyanta hurneralis Ruckes Lutz ' 57'' (5 222 AMNH, FSCA) ; " Coripata 1700m. Yungas La Paz Bol. 1 -XII- 1984 Coll. LE.Pena" (622 USNM) ; " Pte. Mururata Yungas La Paz Bol. 1200-1600rn. 24-26-XII- 1984 Coll. LE.Pena" (455 522 USNM) ; " Chulurnani Yungas La Paz Bol. XII- 1984 Coll. LE.Pena" (2 USNM) ; (a) " BOLIVIA: Dpt. La Paz, Prov. Sud Yungas, 21 km. W. Chulurnani . 4050'. 27-V- 1989. J.E. Eger, coll." (b) "J.E. Eger Collection" (255 422 EGER) ; (a) " BOLIVIA: Dpt. La Paz, Prov. Sud Yungas, Puente Villa . 4300 '. 19-24-V- 1989. J.E. Eger, coll." (b) "J.E. Eger Collection" (255 522 EGER) ; "(SE) Coroico 1800-2100rn. La Paz Bol. 30-XI-2-XII-84 Coll. LE.Pena G." (5 USNM) ; " Rio Coroico 1200m. La Paz Bol. 24-26-XI-84 Coll. LE.Pena" (l055 622 USNM) ; (a) " Bolivia, Coroico 20.12.48 A. Martinez " (b) "CJ Drake Coll. 1956" (5 USNM) ; (a) " Coroico Bolivia " (b) "H G Barber Colin 1950" (5 '? USNM) ; (a) " BOLIVIA, LP., 1190 rn., 1 rni,E. Puente Villa, S.Yungas IV-8-1978 C&L O'Brien" (b) " Thyanta misc 22" (2 ENGL) ; " Circuata-Cajuata 2400 rn. La Paz Bol. 3-5-XIl-84 Coll. LE.Pena" (255 222 USNM) ; " Monteagudo Chuquisaca Bol. 24-XII- 84 Coll. LE.Pena" (422 USNM) ; "(E) Muyuparnpa 1600 rn. Chuquisaca Bol. 21-25-XII-84 Coll. LE.Pena G." (255 322 USNM) ; " Mataral (N) V. Grande Bol. 1800-2000rn. 15-17-XII-1984 Coll. LE.Pena" (2 USNM) ; " Sta. Rosa 1100 rn. (N) Mataral Bol. l 5-XII-84 Coll. LE.Pena" (355 322 USNM) ; " Pto. Camacho (S) Sta. Cruz Bol. 20-XII-84 Coll. LE.Pena" (2 USNM) ; " Cornarapa 1800 rn. Santa Cruz Bol. 14-XII-84 Coll. LE.Pena" (2 USNM) ; " TRES ESTEROS Guanay, Boliv 19/ 25 -Aug-89 leg: LE. Pena" (2 USNM) ; (a) " Rurrenabaque Beni Bolivia WMMMann" (b) " Nov. 1921 " (c) "MULFORD BIOLOGICAL EXPLORATION 1921- 1922" {9 USNM) ; " Coripata " (5 522 MLP), except 12 with (b) "Thyanta, P. DENIER det.'' ( MLP) ; (a) "Ost Bolivien Prov. Lara 750 rn Steinbach S.V." (b) "Z.M. B. Hern." ('? ZMB) ; " caranavi " (2 MLP) ; " Corzuela n 8.1.36 " (2 MLP) ; " Peru, 2400m alt. Dept Cusco Machu Picchu VII, 14-15,1951 sweepingG.H. Dieke" (S' USNM) ; (a) " Macchu Picchu Ruins , Cuzco, Peru March 6 1947 Alt. 9500 ft. " (b) "J. C. Pallister Coll. Donor Frank Johnson" (c) " Thyanta patruelis Stal det. H. Ruckes" (2 AMNH) ; " PERU: Cuzco, Pisac , 3,000m. 15.viii.1971 C. & M. Vardy B.M. 1971- 533" (c BMNH) ; (a) " Abancay , PERU. III-6-51 " (b) "Ross and Michelbacher Collectors'' (222 CAS) ; (a) " Arg. Salta Positos Il. 50 A. Martinez " (b) "CJ Drake Coll. 1956" (5 USNM) ; and (a) "AcSA: 217C ARGENTINA TUCUMAN Cadillal s/Solanurn auriculaturn 15/ 11 /85 ERG" (b) " Thyanta sp. Det. T. J. Henry 1987" (5 USNM) .

Distribution. Southeastern Peru, Bolivia, and northern Argentina ( Map 3).

Comments. In general appearance this species resembles larger specimens of T. patruelis , but it is more closely related to T. brasiliensis . Male specimens can be separated from all other species in the subgenus Agrosoma by the elongate, linear spiculate area on the lateral surface of each pararnere. Male and female specimens can usually be distinguished from T. brasiliensis by the less prominent humeral angles. The only way to reliably separate females of T. boliviensis and T. patruelis is by examining the sperrnatheca of each species. In T. boliviensis , the sclerotized rod is neither swollen subapically nor abruptly narrowed apically as it is in T. patruelis .

Only T. brasiliensis, T. emarginata, T. excavata, and T. hamulata have the sclerotized rod as described above. Thyanta emarginata can be identified by the distinctly excavated basal plates; the remaining three species can be distinguished by the condition of the dilation of the spermatheca. In T. boliviensis this structure is in the form of a single balloon-like structure; in T. brasiliensis it is abruptly narrowed for the distal half; and in T. hamulata it is constricted in the middle and then dilates again, forming a figure 8 shape.

Etymology. Named for the country of the type locality.