Umbilicus Umbilicus, (Salisb.) Dandy Umbilicus Umbilicus, (Guss.) DC.

4. U. rupestris (Salisb.) Dandy View in CoL in Riddelsd., Hedley & Price, Fl. Gloucestershire 611 (1948)

( U. pendulinus DC. , Cotyledon pendulina (DC.) Batt. ).

Stem 20-50 cm, erect. Basal leaves orbicular, peltate, concave above, sinuate-crenate; cauline leaves progressively smaller, mostly reniform, dentate, the uppermost sometimes linear. Bracts usually linear, about equalling the pedicels, but occasionally large and leaf-like. Pedicels 3-9 mm. Flowers 7-10 mm, tubular, usually pendent; raceme fairly dense, sometimes branched at the base, occupying more than half the stem. Sepals ovate, acuminate. Corolla whitish-green or strawcoloured, sometimes tinged with pink; tube about 4 times as long as the ovate, mucronate lobes. Carpels tapered to a fairly long style. S. & W. Europe, northwards to Scotland. Al Az BI Br?Bu Co Cr G a G r H b Hs It Ju Lu Sa Si?Tu. A very variable species; some of the variants resemble 3, 5 and 6 in single characters and have caused much confusion. The most distinct is U. neglectus (Coutinho) Rothm. & P. Silva , Agron. Lusit. 2: 88 (1940), from Portugal, S. Spain and Islas Baléares, which possibly deserves subspecific status. It is a robust plant, with basal leaves often cordate rather than peltate, and a narrow corolla approaching that of 5.

5. U. horizontalis (Guss.) DC. View in CoL , Prodr. 3: 400 (1828)

( Cotyledon horizontalis Guss. ).

Like 4, but with the raceme occupying not more than half the stem; cauline leaves more numerous and crowded, many of them usually linear; flowers narrower and somewhat shorter (c. 7 x 3 mm), subsessile, horizontal; and corolla-lobes triangular, lanceolate, acuminate. 2n=24. Mediterranean region, Bulgaria, Açores. Al Az BI Bu Cr G r Hs It Ju?Sa Si. The plants of the W. Mediterranean region are sometimes separated as U. gaditanus Boiss. , Diagn. Pl. Or. Nov. 1 (6): 58 (1846), and are in some features transitional to 4, but neither their taxonomic status nor their correct name can be ascertained with certainty from the material available. U. intermedius Boiss. , FI. Or. 2: 769 (1872) ( Cotyledon intermedia (Boiss.) Stefanov ), has been recorded for the Balkan peninsula (Bulgaria and Thrace), but its occurrence in Europe is doubtful. It is a plant of S.W. Asia, differing from 4 in its shorter raceme, short pedicels and slightly more pointed corolla-lobes, and from 5 by the usually drooping flowers and corolla with broader tube and lobes.