Dryopteris Adanson

2. Dryopteris Adanson 3

Rhizomes short, erect or ascending, densely covered with broad, soft, often fimbriate scales. Leaves tufted, 1 - to 4-pinnate; veins free. Sori orbicular; indusium reniform.

1 Leaves pinnate with deeply pinnatifid pinnae, or 2-pinnate

2 Leaves10-20 x 2 -3cm,narrowlylanceolate 10. fragrans 2 Leaves normally much larger and broader, lanceolate or oblong

3 Fertile leaves longer and more erect than the sterile leaves; pinnae 10-20 on each side 5. cristata 3 Fertile leaves similar to the sterile leaves; pinnae 20-35 on each side (1-3). filix-mas group 1 Leaves 2-pinnate with pinnatifid pinnules, or 3- to 4-pinnate

4 Leaves 2-pinnate with pinnatifid pinnules; proximal 3 or 4 pinnules on both sides of basal pinna ± equal in length

4. villarii

4 Leaves virtually 3- or 4-pinnate; proximal basiscopic pinnule on basal pinna much longer than the proximal acroscopic pinnule

5 Scales on the petiole concolorous; glands on the indusium sessile or absent

6 Leaves lanceolate or ovate-lanceolate; pinnules ±flat; indusium eglandular 6. carthusiana

6 Leaves triangular-ovate or -lanceolate; pinnules ± concave; indusium with sessile glands on the margin 9. aemula

5 Scales on the petiole usually bicolorous, dark brown or blackish in the centre and pale brown at the margin;

glands on the indusium stalked

7 Proximal basiscopic pinnule of basal pinna nearly always less than half as long as the pinna; spores dark brown with obtuse tubercles 7. dilatata

7 Proximal basiscopic pinnule of basal pinna at least half as long as the pinna; spores pale brown with acute tubercles

8. assiniilis (1-3). D. filix-mas group. Rhizome erect. Fertile leaves similar to sterile leaves, pinnate with deeply pinnatifid pinnae, or 2-pinnate; pinnae 20-35 on each side, teeth not mucronate. 1 Point of insertion of secondary rhachis blackish; pinnules or their ultimate segments usually obliquely truncate, denticulate only at apex 2. borreri 1 Point of insertion of secondary rhachis not blackish; pinnules or their ultimate segments rounded, denticulate, crenate, serrate or lobed all round

2 Large plant with 1 or only a few crowns; usually 3 or more sori on each pinnule; sori 1-5 mm 1. filix-mas 2 Small plant with several crowns; usually 2 sori at most on each pinnule; sori usually less than 1 mm 3. abbreviata