Enica longirostris ( Wiedemann, 1819 )

Enica longirostris ( Wiedemann, 1819) View in CoL

Fig. 10 View Figure 10

Anthrax longirostris Wiedemann, 1819: 11. View in CoL

Cyllenia afra Wiedemann, 1828: 358. View in CoL

Cyllenia pluricellata Macquart, 1855: 104. View in CoL

Taxon depository.

ZooBank: https://zoobank.org/b4e3556d-a8d8-4dd0-924b-7b7f8689dab2.

Diagnosis.

The species is distinguished from other species in the genus by the two crossveins connecting R 1 and R 2 + 3 and by densely setose gena.

Description.

Head: wider than high, creamy white; males holoptic, females dichoptic, vertex and compound eyes at same level; frons (at level of antennal insertion) more or less parallel-sided, female: light brown to reddish-orange, darker than head, without dark spot, males: similar to head, slightly darker than head; frons black setose, with white scales, broadly setose near eyes, extending ventral to antennae; ocellar tubercle slightly raised, dark reddish brown to black, black macrosetose, white or light brown scales posteriorly; occiput whitish – yellow setose, brown setose on occiput spots, white scales antero-dorsally, antero-dorsally with dark brown to black spots, usually oval; gena creamy white, densely white setose.

Proboscis and maxillary palpus: proboscis straight, black; labella only forming distal tip of proboscis, apically rounded, minute black setose; maxillary palpus not extending beyond oral cavity, light brown to brown, brown setose or light yellowish-brown setose.

Antenna: dark brown; scape creamy white, approximately as long as pedicel, variable setation color: black setose, white setose, or white scales may be present, setae extending to tip of pedicel; pedicel reddish brown or brown, short black setose dorsally; postpedicel black or dark brown, tapering distally, longer than scape and pedicel combined, white or light yellow scales dorsally; stylus reduced, only apical ‘ seta-like’ sensory element present, situated sub – apically in cavity on postpedicel.

Thorax: light orangish brown or reddish brown and light orangish brown; scutum predominantly black, reddish on margins, light to dark brown scales, two submedial longitudinal stripes with lighter scales; setation: acr setae present, black, dc setae present, black; prosternum silvery grey pubescence; proepisternum silvery grey pubescence, long yellowish-white setose, white scales; antepronotum silvery grey pubescence, dull white scales anteriorly; postpronotum silvery grey pubescence, asetose, with medial dark brown to black stripe; postpronotal lobe silvery grey pubescence, white scales, white and black setose; pleuron silvery grey pubescence; proepimeron silvery grey pubescence, asetose; anepisternum silvery grey pubescence, white scales ventrally, black macrosetose dorsally; anepimeron silvery grey pubescence, asetose or white scales; katepisternum silvery grey pubescence, white scales dorsally; laterotergite silvery grey pubescence, asetose; meron + metepisternum silvery grey pubescence, asetose or white scales dorsally; metepimeron silvery grey pubescence, asetose or white setose dorsally; scutellum red, sometimes with anterior or posterior black spots, apubescent, ds sctl setae black macrosetose, ap sctl setae comprised of light brown and white scales.

Leg: light orangish brown, covered in scales, some regular setae present; pro coxa light orangish brown, silvery grey pubescence, white scales, thick black setose; pro femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; pro tibia light brown, raised short and fine dark brown setose ventrally, large black setose dorsally; mes coxa light orangish brown, white pubescence, white scales, thick black setose; mes femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; mes tibia light brown, dark brown scales, long black setose, distal tip with long black setae; met coxa light orangish brown, white pubescent, white scales, thick black setose, anteriorly without any protuberance; met trochanter short black setose ventrally, white scales dorsally, without protuberance; met femur light orangish brown, white scales ventrally, brown scales dorsally, few short black setae; met tibia light brown, dark brown scales, large black setose, distal tip with long black macrosetae of varying sizes; proximal pro, mes, and met tarsomeres longer than following 2 tarsomeres combined, proximal met tarsomere as wide as following tarsomeres; pro tarsomere with dark brown scales dorsally, black setose ventrally; mes tarsomere with dark brown scales dorsally, black setose ventrally; met tarsomere with dark brown scales dorsally, black setose ventrally; claw smoothly arched distally.

Wing: (6.1 –) 9.1–9.8 (– 11.4) mm, microtrichia absent; membrane color pattern males: predominantly brown colored except for white tinted base of wing, supernumerary crossveins between R 1 and R 2 + 3, oblique rectangular area between CuA and R 1, crossveins r-r and r-m, and base of cell m 2, females: membrane dark brown colored except for white tinted base of wing, supernumerary crossveins between R 1 and R 2 + 3, oblique rectangular area between CuA and R 1, crossveins r-r and r-m, and base of cell m 2; costal vein dark brown short setose, whitish clear setose proximally; R 2 + 3 distally distinctly arching anteriorly, r 1 open, divided by 2 supernumerary crossveins; R 4 terminating anterior to wing apex, distinctly arching anteriorly; cell r 4 open, proximally R 4 and R 5 parallel, R 4 strongly diverging anteriorly in distal 2 / 3; R 5 terminating posterior to wing apex; r 5 open; M 1 terminating posterior to wing apex; cell d closed by base of M 2 and m-m, r-m situated in distal ¼; m 3 open; cua open; alula greatly reduced, nearly straight wing margin; haltere stem dark brown setose.

Abdomen: shape ovate, black, predominantly black anteriorly and red posteriorly or reddish brown; tergites smooth, setae with small sockets only; T 1 reddish yellow to light brown, reddish brown, or predominantly black, no red or a very thin line of red posteriorly, light yellowish-brown to light whitish brown setose, scales or snow – white setose, with scales; T 2 – T 7 predominantly reddish-orange or predominantly black or predominantly dark reddish brown, light brown setose, white scales, rarely brown to light yellowish-brown setose, white scales or dark brown setose, white scales, white scales on medial spots on T 2–8 and posterior margins of T 2 and T 4, rarely scales uniformly color throughout or light brown setose, 2 spots with dark scales on T 2 and T 3; S 1–8 predominantly black with reddish brown, yellow to white scales, thin black setose and long white setose or brown scales, thin black setose and long white setose; lateral margin lacking setae, rarely sparse black setae on T 4–8, male T 8 white scales, black setose.

Terminalia: Male terminalia drawings in Hesse (1956, p 32): https://www.biodiversitylibrary.org/page/40844167.

Other material.

South Africa – Eastern Cape • 1 ♀ Alicedale , 2 km E; 33°19'07"S, 026°06'12"E; 310 m a. s. l.; 23 Oct. 2004; Londt, Jason leg.; Dry rocky hillside with succulent vegetation; NMSA -DIP-93654 , NMSA GoogleMaps • 1 ♂ Alicedale , 2 km E; 33°19'07"S, 026°06'12"E; 310 m a. s. l.; 23 Oct. 2004; Londt, Jason leg.; Dry rocky hillside with succulent vegetation; NMSA -DIP-93655 , NMSA GoogleMaps • 1 ♀ Alicedale , 2 km E; 33°19'07"S, 026°06'12"E; 310 m a. s. l.; 23 Oct. 2004; Londt, Jason leg.; Dry rocky hillside with succulent vegetation; NMSA -DIP-93656 , NMSA GoogleMaps • 1 ♀ 3 ♂ 1? Groendal, Uitenhage (= kariega); 33°43'13"S, 025°18'55"E; 28 Oct. 1938; SAM Museum Staff leg.; SAM-DIP-A 002141 , SAMC GoogleMaps • 1 ♀ Willowmore ; 33°17'00"S, 023°29'00"E; 25 Oct. 1918; Brauns, H. leg.; NMSA -DIP-054408 , NMSA GoogleMaps • 1 ♀ Willowmore ; 33°17'00"S, 023°29'00"E; 01 Nov. 1919; Brauns, H. leg.; NMSA -DIP-054687 , NMSA GoogleMaps • 1 ♀ Willowmore ; 33°17'00"S, 023°29'00"E; 01 Nov. 1919; Brauns, H. leg.; NMSA -DIP-89485 , NMSA GoogleMaps • 1 ♂ Willowmore ; 33°17'00"S, 023°29'00"E; 15 Oct. 1921; Brauns, H. leg.; NMSA -DIP-056255 , NMSA GoogleMaps • 1 ♂ Willowmore ; 33°17'00"S, 023°29'00"E; 15 Oct. 1921; Brauns, H. leg.; NMSA -DIP-89530 , NMSA GoogleMaps • 1 ♀ Willowmore ; 33°17'00"S, 023°29'00"E; Dec. 1922; Brauns, H. leg.; NMSA -DIP-89522 , NMSA GoogleMaps • 1 ♂ Willowmore ; 33°17'00"S, 023°29'00"E; Dec. 1922; Brauns, H. leg.; NMSA -DIP-89523 , NMSA GoogleMaps

South Africa – Northern Cape • 1 ♀ Bowesdorp (= Bowes Dorp), Namaqualand ; 30°09'03"S, 017°55'55"E; Nov. 1931; SAM Museum Staff leg.; SAM-DIP-A 002167 , SAMC GoogleMaps • 2 ♀ Hester Malan N. R. (= Goegap Nature Reserve), Namaqualand ; 29°40'00"S, 018°02'00"E; 16 Oct. 1985; Struck, M. leg.; SAM-DIP-A 004968 , SAMC GoogleMaps • 1 ♀ Kamieskroon Namaqualaud ; 30°12'00"S, 017°56'00"E; Nov. 1936; SAM Museum Staff leg.; SAM-DIP-A 002143 , SAMC GoogleMaps • 1 ♀ Kamieskroon , 10 km E; 30°11'39"S, 018°00'10"E; 630 m a. s. l.; 17 Oct. 1977; Miller, R. M. leg.; NMSA -DIP-049597 , NMSA GoogleMaps • 1 ♂ Nieuwoudtville ; 31°23'00"S, 019°06'00"E; Aug. 1928; Brauns, H. leg.; NMSA -DIP-056002 , NMSA GoogleMaps • 1 ♂ Nieuwoudtville, 1 km W, caravan park and area ; 31°23'00"S, 019°06'00"E; 800 m a. s. l.; 04 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92784 , NMSA GoogleMaps • 1 ♀ Nieuwoudtville, 1 km W, caravan park and area ; 31°23'00"S, 019°06'00"E; 800 m a. s. l.; 04 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92785 , NMSA GoogleMaps • 1 ♀ Nieuwoudtville, 9 km W, Summit Vanrhyn’s Pass ; 31°22'00"S, 019°01'00"E; 700 m a. s. l.; 04 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92829 , NMSA GoogleMaps • 1 ♂ Outiep Garies Namaqualand (= Namakwaland ); 30°33'27"S, 017°59'34"E; Sep. 1953; Toit, J. leg.; SAM-DIP-A 002145 , SAMC GoogleMaps • 1 ♀ Springbok, 8 km NW, road to Nababeep ; 29°37'16"S, 017°49'59"E; 985 m a. s. l.; 30 Sep. 2009; Londt, Jason, Dikow, Torsten leg.; thick woody fynbos; NMSA -DIP-92350 , NMSA GoogleMaps • 1 ♂ Springbok, 8 km NW, road to Nababeep ; 29°37'16"S, 017°49'59"E; 985 m a. s. l.; 30 Sep. 2009; Londt, Jason, Dikow, Torsten leg.; thick woody fynbos; NMSA -DIP-92351 , NMSA GoogleMaps

South Africa – Western Cape • 1 ♂ Brenton on Sea ; 34°04'20"S, 023°01'07"E; 10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; dune and hillside vegetation; NMSA -DIP-050742 , NMSA GoogleMaps • 1 ♂ Brenton on Sea ; 34°04'20"S, 023°01'07"E; 10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; dune and hillside vegetation; NMSA -DIP-89541 , NMSA GoogleMaps • 1 ♀ Brenton on Sea ; 34°04'20"S, 023°01'07"E; 10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; dune and hillside vegetation; NMSA -DIP-89542 , NMSA GoogleMaps • 1 ♀ Cape Town ; 33°59'00"S, 018°26'00"E; 1874; Trimen, R. leg.; SAM-DIP-A 002169 , SAMC GoogleMaps • 1 ♂ Cape Town ; 33°59'00"S, 018°26'00"E; 1915; Péringuey, L. leg.; SAM-DIP-A 002142 , SAMC GoogleMaps • 1 ♀ Ceres, upper sources of Olifants River ; 33°12'29"S, 019°14'49"E; Oct. 1949; SAM Museum Staff leg.; SAM-DIP-A 002118 , SAMC GoogleMaps • 3 ♀ 3 ♂ Ceres, upper sources of Olifants River ; 33°12'29"S, 019°14'49"E; Dec. 1949; SAM Museum Staff leg.; SAM-DIP-A 002118 , SAMC GoogleMaps • 1 ♀ 1 ♂ Ceres, upper sources of Olifants River ; 33°12'29"S, 019°14'49"E; Dec. 1949; SAM Museum Staff leg.; SAM-DIP-A 002118 , SAMC GoogleMaps • 1 ♂ Ceres, upper sources of Olifants River ; 33°12'29"S, 019°14'49"E; Dec. 1949; SAM Museum Staff leg.; SAM-DIP-A 002118 , SAMC GoogleMaps • 1 ♂ Ceres, upper sources of Olifants River ; 33°12'29"S, 019°14'49"E; Dec. 1949; SAM Museum Staff leg.; SAM-DIP-A 002118 , SAMC GoogleMaps • 1 ♀ Citrusdale , 14 km NNW; 32°31'00"S, 018°58'00"E; 300 m a. s. l.; 01 Nov. 1991; Londt, Jason leg.; woody plants, sandy; NMSA -DIP-92878 , NMSA GoogleMaps • 1 ♂ Citrusdale , 14 km NNW; 32°31'00"S, 018°58'00"E; 300 m a. s. l.; 01 Nov. 1991; Londt, Jason leg.; woody plants, sandy; NMSA -DIP-92879 , NMSA GoogleMaps • 1 ♀ Citrusdale , 14 km NNW; 32°31'00"S, 018°58'00"E; 300 m a. s. l.; 01 Nov. 1991; Londt, Jason leg.; woody plants, sandy; NMSA -DIP-92880 , NMSA GoogleMaps • 1 ♂ Citrusdale , 14 km NNW; 32°31'00"S, 018°58'00"E; 300 m a. s. l.; 01 Nov. 1991; Londt, Jason leg.; woody plants, sandy; NMSA -DIP-92881 , NMSA GoogleMaps • 1 ♀ Clanwilliam , 4 m. S; 32°13'18"S, 018°51'08"E; Sep. 1961; SAM Museum Staff leg.; SAM-DIP-A 002168 , SAMC GoogleMaps • 1 ♀ coast near Swartklip , 20 km SW Stellenbosch; 34°04'00"S, 018°43'00"E; 0 m a. s. l.; 30 Oct. 1991; Londt, Jason leg.; NMSA -DIP-92687 , NMSA GoogleMaps • 1 ♀ coast near Swartklip , 20 km SW Stellenbosch; 34°04'00"S, 018°43'00"E; 0 m a. s. l.; 30 Oct. 1991; Londt, Jason leg.; NMSA -DIP-92836 , NMSA GoogleMaps • 1 ♂ coast near Swartklip , 20 km SW Stellenbosch; 34°04'00"S, 018°43'00"E; 0 m a. s. l.; 30 Oct. 1991; Londt, Jason leg.; NMSA -DIP-92837 , NMSA GoogleMaps • 1 ♂ coast near Swartklip , 20 km SW Stellenbosch; 34°04'00"S, 018°43'00"E; 0 m a. s. l.; 30 Oct. 1991; Londt, Jason leg.; NMSA -DIP-92841 , NMSA GoogleMaps • 1 ♂ Darling Wild Flower Reserve ; 33°23'14"S, 018°22'56"E; 180 m a. s. l.; 13 Oct. 2006; Londt, Jason leg.; Mixed fynbos, sandy area; NMSA -DIP-92367 , NMSA GoogleMaps • 1 ♂ Darling Wild Flower Reserve ; 33°23'14"S, 018°22'56"E; 180 m a. s. l.; 13 Oct. 2006; Londt, Jason leg.; Mixed fynbos, sandy area; NMSA -DIP-92368 , NMSA GoogleMaps • 1 ♀ Darling Wild Flower Reserve ; 33°23'14"S, 018°22'56"E; 180 m a. s. l.; 13 Oct. 2006; Londt, Jason leg.; Mixed fynbos, sandy area; NMSA -DIP-92369 , NMSA GoogleMaps • 1 ♀ Darling Wild Flower Reserve ; 33°23'14"S, 018°22'56"E; 180 m a. s. l.; 13 Oct. 2006; Londt, Jason leg.; Mixed fynbos, sandy area; NMSA -DIP-92370 , NMSA GoogleMaps • 1 ♂ Darling Wild Flower Reserve ; 33°23'14"S, 018°22'56"E; 180 m a. s. l.; 13 Oct. 2006; Londt, Jason leg.; Mixed fynbos, sandy area; NMSA -DIP-92371 , NMSA GoogleMaps • 1 ♀ Darling , 3 km S; 33°24'21"S, 018°22'51"E; 28 Sep. 1979; Londt, Jason leg.; well veget. hillside above wheatlands; NMSA -DIP-050598 , NMSA GoogleMaps • 1 ♀ Darling , 3 km S; 33°24'21"S, 018°22'51"E; 28 Sep. 1979; Londt, Jason leg.; well, veget. hillside above wheatlands; NMSA -DIP-89545 , NMSA GoogleMaps • 3 ♂ Eland’s Bay, Leipoldtville ; 32°18'44"S, 018°20'29"E; Oct. 1947; SAM Museum Staff leg.; SAM-DIP-A 002134 , SAMC GoogleMaps • 1 ♂ Elandsbaai , 2.5 mi. S; 32°21'09"S, 018°19'32"E; 30 m a. s. l.; 16 Sep. 1972; Irwin, M. E., Irwin, B. J. leg.; coastal sand dunes; NMSA -DIP-050959 , NMSA GoogleMaps • 1 ♀ Elephant Leisure Resort , 8 km S of Citrusdal; 32°40'33"S, 019°00'34"E; 250 m a. s. l.; 11 Oct. 2014; Reavell, P. leg.; NMSA -DIP-192817 , NMSA GoogleMaps • 1 ♂ Fish Hoek., Cape Peninsula ; 34°07'55"S, 018°26'13"E; Oct. 1931; Munro, H. leg.; NMSA -DIP-049316 , NMSA GoogleMaps • 1 ♀ Fish Hoek., Cape Peninsula ; 34°07'55"S, 018°26'13"E; Oct. 1931; Munro, H. leg.; NMSA -DIP-89484 , NMSA GoogleMaps • 1 ♂ Gaukamma Nature Reserve ; 34°02'00"S, 022°57'00"E; 25 Dec. 1988; Londt, Jason leg.; Macchia sandy ground and shrubs; NMSA -DIP-056053 , NMSA GoogleMaps • 1 ♀ Gaukamma Nature Reserve ; 34°02'00"S, 022°57'00"E; 25 Dec. 1988; Londt, Jason leg.; Macchia sandy ground and shrubs; NMSA -DIP-89521 , NMSA GoogleMaps • 1 ♂ Gifberg Pass summit ; 31°46'00"S, 018°45'00"E; 690 m a. s. l.; 16 Nov. 1986; Londt, Jason, Quickelberge, C. leg.; rocky macchia; NMSA -DIP-053209 , NMSA GoogleMaps • 1 ♂ Gifberg Pass , 21 km south Vanrhynsdorp; 31°46'21"S, 018°45'59"E; 540 m a. s. l.; 08 Sep. 2002; Londt, Jason leg.; montane macchia; NMSA -DIP-84612 , NMSA GoogleMaps • 1 ♂ Gifberg Pass , 21 km south Vanrhynsdorp; 31°46'21"S, 018°45'59"E; 540 m a. s. l.; 08 Sep. 2002; Londt, Jason leg.; montane macchia; NMSA -DIP-84613 , NMSA GoogleMaps • 1 ♀ Gifberg , flat summit, 24 km S Vanrhysdorp; 31°46'00"S, 018°48'00"E; 600 m a. s. l.; 03 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92780 , NMSA GoogleMaps • 1 ♂ Gifberg , flat summit, 24 km S Vanrhysdorp; 31°46'00"S, 018°48'00"E; 600 m a. s. l.; 03 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92781 , NMSA GoogleMaps • 1 ♀ 1 ♂ Gifberg , flat summit, 24 km S Vanrhysdorp; 31°46'00"S, 018°48'00"E; 600 m a. s. l.; 03 Nov. 1991; Londt, Jason leg.; NMSA -DIP-92782 , NMSA GoogleMaps • 1 ♂ Goukamma Nature Reserve, along Bush Pig trail ; 34°03'41"S, 022°56'08"E; 29 m a. s. l.; 09 Dec. 2015; collected p. m. (noon – 15: 00); Dikow, Torsten leg.; fynbos covered sand dune; USNMENT 01115015 , USNM GoogleMaps • 1 ♂ Goukamma Nature Reserve, along Bush Pig trail ; 34°03'41"S, 022°56'08"E; 29 m a. s. l.; 09 Dec. 2015; collected p. m. (noon – 15: 00); Dikow, Torsten leg.; fynbos covered sand dune; USNMENT 01115042 , USNM GoogleMaps • 1 ♂ Goukamma Nature Reserve, along Bush Pig trail ; 34°03'41"S, 022°56'08"E; 29 m a. s. l.; 09 Dec. 2015; collected p. m. (noon – 15: 00); Dikow, Torsten leg.; fynbos covered sand dune; USNMENT 01115157 , USNM GoogleMaps • 1 ♀ Goukamma Nature Reserve, along Bush Pig trail ; 34°03'41"S, 022°56'08"E; 29 m a. s. l.; 09 Dec. 2015; collected p. m. (noon – 15: 00); Dikow, Torsten leg.; fynbos covered sand dune; USNMENT 01115166 , USNM GoogleMaps • 1 ♂ Goukamma Nature Reserve, along Bush Pig trail ; 34°03'41"S, 022°56'08"E; 29 m a. s. l.; 09 Dec. 2015; collected p. m. (noon – 15: 00); Dikow, Torsten leg.; fynbos covered sand dune; USNMENT 01115171 , USNM GoogleMaps • 1 ♂ Goukamma Nature Reserve, along Bush Pig trail ; 34°03'41"S, 022°56'08"E; 29 m a. s. l.; 09 Dec. 2015; collected p. m. (noon – 15: 00); Dikow, Torsten leg.; fynbos covered sand dune; USNMENT 01115221 , USNM GoogleMaps • 1 ♂ Hout Bay ; 34°02'55"S, 018°19'55"E; Nov. 1943; leg.; SAM-DIP-A 016116 , SAMC GoogleMaps • 1 ♂ Hout Bay ; 34°02'55"S, 018°19'55"E; Nov. 1943; leg.; SAM-DIP-A 016117 , SAMC GoogleMaps • 1 ♂ Kagga Kamma Nature Reserve ; 32°45'09"S, 019°34'13"E; 1075 m a. s. l.; 22–23 Nov. 2008; Londt, Jason leg.; Sandy area with tall fynbos near houses; NMSA -DIP-92985 , NMSA GoogleMaps • 1 ♂ Kagga Kamma Nature Reserve ; 32°45'09"S, 019°34'13"E; 1075 m a. s. l.; 22–23 Nov. 2008; Londt, Jason leg.; Sandy area with tall fynbos near houses; NMSA -DIP-92986 , NMSA GoogleMaps • 1 ♂ Klaver (= Klawer ); 31°46'29"S, 018°37'07"E; 19 Sep. 1917; Roberts, A. leg.; NMSA -DIP-048223 , NMSA GoogleMaps • 2 ♀ 2 ♂ Kleinmond ; 34°20'43"S, 019°00'52"E; Jan. 1937; SAM Museum Staff leg.; SAM-DIP-A 002135 , SAMC GoogleMaps • 1 ♀ Knersvlakte, North of Van Rhynsdorp , SW cape; 31°15'00"S, 018°45'00"E; 06–09 Oct. 1964; Stuckenberg, Brian, Stuckenberg, P. leg.; NMSA -DIP-89508 , NMSA GoogleMaps • 1 ♂ Knersvlakte, North of Van Rhynsdorp , SW cape; 31°15'00"S, 018°45'00"E; 06–09 Oct. 1964; Stuckenberg, Brian, Stuckenberg, P. leg.; NMSA -DIP-89509 , NMSA GoogleMaps • 1 ♂ Knersvlakte, North of Van Rhynsdorp , SW cape; 31°15'00"S, 018°45'00"E; 06–09 Oct. 1964; Stuckenberg, Brian, Stuckenberg, P. leg.; NMSA -DIP-89513 , NMSA GoogleMaps • 1 ♀ Knysna Heads , N side; 34°04'25"S, 023°03'55"E; 09–10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; open hillside grass; NMSA -DIP-055992 , NMSA GoogleMaps • 1 ♂ Knysna Heads , N side; 34°04'25"S, 023°03'55"E; 09–10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; open hillside grass; NMSA -DIP-89514 , NMSA GoogleMaps • 1 ♂ Knysna Heads , N side; 34°04'25"S, 023°03'55"E; 09–10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; open hillside grass; NMSA -DIP-89543 , NMSA GoogleMaps • 1 ♂ Knysna Heads , N side; 34°04'25"S, 023°03'55"E; 09–10 Dec. 1979; Londt, Jason, Stuckenberg, Brian leg.; open hillside grass; NMSA -DIP-89544 , NMSA GoogleMaps • 1 ♂ Knysna, Keurbooms River ; 33°53'12"S, 23°14'30"E; Jan. 1931; Barnard, K. leg.; SAM-DIP-A 002144 , SAMC GoogleMaps • 1 ♂ Kommetjie ; 34°09'00"S, 018°20'00"E; 30 m a. s. l.; 01 Oct. 1993; Londt, Jason leg.; Macchia, sandy area, hillside; NMSA -DIP-95244 , NMSA GoogleMaps • 1 ♂ Kommetjie ; 34°09'00"S, 018°20'00"E; 30 m a. s. l.; 01 Oct. 1993; Londt, Jason leg.; Macchia, sandy area, hillside; NMSA -DIP-95245 , NMSA GoogleMaps • 1 ♀ Kommetjie , hillside; 34°09'00"S, 018°20'00"E; 30 m a. s. l.; 01 Oct. 1993; Londt, Jason leg.; Macchia, sandy area; NMSA -DIP-95243 , NMSA GoogleMaps • 1 ♀ Laaiplek ; 32°47'10"S, 018°09'40"E; 09 Oct. 1977; Miller, R. M. leg.; NMSA -DIP-049611 , NMSA GoogleMaps • 1 ♀ Laaiplek ; 32°47'10"S, 018°09'40"E; 09 Oct. 1977; Miller, R. M. leg.; NMSA -DIP-89517 , NMSA GoogleMaps • 1 ♂ Laaiplek ; 32°47'10"S, 018°09'40"E; 09 Oct. 1977; Miller, R. M. leg.; NMSA -DIP-89518 , NMSA GoogleMaps • 1 ♂ Lamberts B. P. C. (= Lambert’s Bay ); 32°05'24"S, 018°18'47"E; 20 Nov. 1917; Roberts, A. leg.; NMSA -DIP-048245 , NMSA GoogleMaps • 3 ♂ Moordenaars Karoo Swanepoel ; 32°58'02"S, 020°48'56"E; Oct. 1952; SAM Museum Staff leg.; SAM-DIP-A 002133 , SAMC GoogleMaps • 1 ♀ Muizenberg ; 34°05'24"S, 018°29'45"E; 27 Dec. 1934; van Son, G. leg.; NMSA -DIP-051536 , NMSA GoogleMaps • 1 ♂ Muizenberg (Cape Town), Macchia slopes above ; 34°05'24"S, 018°29'45"E; 28 Nov. 1981; Stuckenberg, Brian leg.; macchia slopes; NMSA -DIP-048697 , NMSA GoogleMaps • 1 ♂ Muizenberg (Cape Town), Macchia slopes above ; 34°05'24"S, 018°29'45"E; 28 Nov. 1981; Stuckenberg, Brian leg.; macchia slopes; NMSA -DIP-89515 , NMSA GoogleMaps • 1 ♂ Nuwekloof , 3 km E Gouda; 33°19'00"S, 019°06'00"E; 120 m a. s. l.; 05 Oct. 1993; Londt, Jason leg.; Old road and riverbanks; NMSA -DIP-93299 , NMSA GoogleMaps • 1 ♂ Nuwekloof , 3 km E Gouda; 33°19'00"S, 019°06'00"E; 120 m a. s. l.; 05 Oct. 1993; Londt, Jason leg.; Old road and riverbanks; NMSA -DIP-93300 , NMSA GoogleMaps • 3 ♀ Olifants River between Citrusdal and Clanwilliam ; 32°26'43"S, 018°57'31"E; Oct. 1931; SAM Museum Staff leg.; SAM-DIP-A 002158 , SAMC GoogleMaps • 1 ♀ 5 ♂ Paleisheuwel ; 32°28'25"S, 018°43'19"E; Nov. 1949; SAM Museum Staff leg.; SAM-DIP-A 002125 , SAMC GoogleMaps • 1 ♀ Paleisheuwel ; 32°28'25"S, 018°43'19"E; Nov. 1949; SAM Museum Staff leg.; SAM-DIP-A 002125 , SAMC GoogleMaps • 1 ♂ Papendrop (= Papendorp) Olifaunts River ; 31°41'55"S, 018°12'34"E; Oct. 1950; SAM Museum Staff leg.; SAM-DIP-A 002148 , SAMC GoogleMaps • 9 ♀ 18 ♂ Pearly Beach, Bredarsdorp Div. ; 34°40'09"S, 019°30'00"E; 01 Dec. 1938; SAM Museum Staff leg.; SAM-DIP-A 016213 , SAMC GoogleMaps • 1 ♀ Pearly Beach, Bredarsdorp Div. ; 34°40'09"S, 019°30'00"E; 01–13 Dec. 1938; SAM Museum Staff leg.; SAM-DIP-A 016206 , SAMC GoogleMaps • 1 ♂ Pearly Beach, Bredarsdorp Div. ; 34°40'09"S, 019°30'00"E; 01–13 Dec. 1938; SAM Museum Staff leg.; SAM-DIP-A 016207 , SAMC GoogleMaps • 2 ♂ Pearly Beach, Bredarsdorp Div. ; 34°40'09"S, 019°30'00"E; 01–13 Dec. 1938; SAM Museum Staff leg.; SAM-DIP-A 016213 , SAMC GoogleMaps • 1 ♂ Pearly Beach, Bredarsdorp Div. ; 34°40'09"S, 019°30'00"E; Dec. 1958; SAM Museum Staff leg.; SAM-DIP-A 002147 , SAMC GoogleMaps • 1 ♂ Plettenberg ; 34°03'00"S, 023°22'00"E; Jan. 1952; leg.; SAM-DIP-A 002149 , SAMC GoogleMaps • 1 ♂ Polsuesl (Pol- ---- l) R.; 34°22'07"S, 018°52'30"E; Dec. 1932; Wood, H. G. leg.; SAM-DIP-A 002137 , SAMC GoogleMaps • 1 ♂ Rocherpan Nature Reserve Trail ; 32°36'20"S, 018°18'24"E; 11 m a. s. l.; 03 Oct. 2019; Cabrero, A., Deschodt, C. leg.; open areas near pan;, Coll. Cabrero GoogleMaps • 1 ♂ Saldanha Bay ( Hopefield crossed out ); 33°00'23"S, 017°56'08"E; Sep. 1960; SAM Museum Staff leg.; SAM-DIP-A 002146 , SAMC GoogleMaps • 1 ♀ Silvermine Nature Reserve ; 34°08'00"S, 018°26'00"E; 22 Dec. 1988; Londt, Jason leg.; Macchia sandy ground and rocks; NMSA -DIP-056051 , NMSA GoogleMaps • 1 ♀ Silvermine Nature Reserve ; 34°05'00"S, 018°25'00"E; 442 m a. s. l.; 16 Dec. 1995; Londt, Jason leg.; NMSA -DIP-95494 , NMSA GoogleMaps • 1 ♀ Silvermine Nature Reserve ; 34°05'00"S, 018°25'00"E; 442 m a. s. l.; 16 Dec. 1995; Londt, Jason leg.; NMSA -DIP-95495 , NMSA GoogleMaps • 1 ♂ Silvermine Nature Reserve ; 34°05'00"S, 018°25'00"E; 442 m a. s. l.; 16 Dec. 1995; Londt, Jason leg.; NMSA -DIP-95496 , NMSA GoogleMaps • 1 ♂ Silvermine Nature Reserve ; 34°05'00"S, 018°25'00"E; 442 m a. s. l.; 16 Dec. 1995; Londt, Jason leg.; NMSA -DIP-95497 , NMSA GoogleMaps • 1 ♀ Silvermine Nature Reserve ; 34°05'00"S, 018°25'00"E; 442 m a. s. l.; 16 Dec. 1995; Londt, Jason leg.; NMSA -DIP-95498 , NMSA GoogleMaps • 1 ♀ Silvermine Nature Reserve ; 34°05'00"S, 018°25'00"E; 442 m a. s. l.; 16 Dec. 1995; Londt, Jason leg.; NMSA -DIP-95502 , NMSA GoogleMaps • 1 ♂ Somerset Strand ; 34°04'52"S, 018°48'52"E; 08 Nov. 1926; Brauns, H. leg.; NMSA -DIP-89527 , NMSA GoogleMaps • 1 ♂ Somerset West Strand ; 34°04'52"S, 018°48'52"E; Oct. 1925; Brauns, H. leg.; NMSA -DIP-054598 , NMSA GoogleMaps • 1 ♂ Somerset West Strand ; 34°04'52"S, 018°48'52"E; 25 Nov. 1925; Brauns, H. leg.; NMSA -DIP-054599 , NMSA GoogleMaps • 1 ♂ Somerset West Strand ; 34°04'52"S, 018°48'52"E; 25 Nov. 1925; Brauns, H. leg.; NMSA -DIP-89494 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 1888; leg.; SAM-DIP-A 002163 , SAMC GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 15 Oct. 1916; Brauns, H. leg.; NMSA -DIP-056227 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 15 Oct. 1916; Brauns, H. leg.; NMSA -DIP-056227 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 15 Oct. 1916; Brauns, H. leg.; NMSA -DIP-89486 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 15 Oct. 1916; Brauns, H. leg.; NMSA -DIP-89498 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; Oct. 1925; Brauns, H. leg.; NMSA -DIP-89487 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; Oct. 1925; Brauns, H. leg.; NMSA -DIP-89495 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 01 Dec. 1925; Brauns, H. leg.; NMSA -DIP-056220 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 05 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056251 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 10 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056252 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 12 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056253 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 14 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89529 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 15 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056247 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 20 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056228 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 20 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056257 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 20 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89496 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 20 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89526 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 20 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89548 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 21 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056258 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 22 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056259 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 22 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89524 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 22 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89528 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 22 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89552 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 25 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056205 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 25 Oct. 1926; Brauns, H. leg.; NMSA -DIP-056260 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 25 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89546 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 25 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89547 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 25 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89553 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 25 Oct. 1926; Brauns, H. leg.; NMSA -DIP-98525 , NMSA GoogleMaps • 1 ♀ Stellenbosch ; 33°56'00"S, 018°51'00"E; 30 Oct. 1926; Brauns, H. leg.; NMSA -DIP-89549 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 04 Nov. 1926; Brauns, H. leg.; NMSA -DIP-89531 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 04 Nov. 1926; Brauns, H. leg.; NMSA -DIP-89532 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 04 Nov. 1926; Brauns, H. leg.; NMSA -DIP-89550 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 04 Nov. 1926; Brauns, H. leg.; NMSA -DIP-89551 , NMSA GoogleMaps • 1 ♂ Stellenbosch ; 33°56'00"S, 018°51'00"E; 15 Nov. 1926; Brauns, H. leg.; NMSA -DIP-056239 , NMSA GoogleMaps • 1 ♀ Table Mountain National Park ; 34°14'27"S, 018°23'10"E; 20 m a. s. l.; 16 Oct. 2006; Londt, Jason leg.; sandy hillside bynbos area with flowers; NMSA -DIP-106783 , NMSA GoogleMaps • 1 ♂ Table Mountain National Park, Cape of Good Hope area ; 34°20'42"S, 018°27'46"E; 20 m a. s. l.; 10 Oct. 2006; Londt, Jason leg.; coastal fynbos; NMSA -DIP-106781 , NMSA GoogleMaps • 1 ♂ Table Mountain National Park, Olifantsbos ; 34°14'27"S, 018°23'10"E; 20 m a. s. l.; 16 Oct. 2006; Londt, Jason leg.; Sandy hillside fynbos area with flowers; NMSA -DIP-106782 , SAMC GoogleMaps • 1 ♂ Table Mountain National Park, Olifantsbos ; 34°14'27"S, 018°23'10"E; 20 m a. s. l.; 16 Oct. 2006; Londt, Jason leg.; Sandy hillside fynbos area with flowers; NMSA -DIP-106784 , NMSA GoogleMaps • 1 ♀ Table Mountain National Park, Olifantsbos ; 34°14'27"S, 018°23'10"E; 20 m a. s. l.; 16 Oct. 2006; Londt, Jason leg.; Sandy hillside fynbos area with flowers; NMSA -DIP-106785 , NMSA GoogleMaps • 1 ♂ Table Mountain National Park, Olifantsbos ; 34°14'27"S, 018°23'10"E; 20 m a. s. l.; 16 Oct. 2006; Londt, Jason leg.; Sandy hillside fynbos area with flowers; NMSA -DIP-106786 , NMSA GoogleMaps • 1 ♂ Table Mountain National Park, Olifantsbos ; 34°14'27"S, 018°23'10"E; 20 m a. s. l.; 16 Oct. 2006; Londt, Jason leg.; Sandy hillside fynbos area with flowers; NMSA -DIP-106787 , NMSA GoogleMaps • 2 ♂ Wit River (= Witrivier) Valley, Bains Kloof (= Bainskloof Pass); 33°34'12"S, 019°08'20"E; Dec. 1949; SAM Museum Staff leg.; SAM-DIP-A 002132 , SAMC GoogleMaps • 1 ♂ Wit River (= Witrivier) Valley, Bains Kloof (= Bainskloof Pass); 33°34'12"S, 019°08'20"E; Dec. 1949; SAM Museum Staff leg.; SAM-DIP-A 002132 , SAMC GoogleMaps • 1 ♀ Ysterfontein (= Yzerfontein ); 33°20'58"S, 018°09'15"E; 20 Oct. 1964; Stuckenberg, Brian leg.; NMSA -DIP-048768 , NMSA GoogleMaps • 1 ♀ Ceres, upper sources of Olifants River ; 33°12'29"S, 019°14'49"E; Dec. 1949; SAM Museum Staff leg.; SAM-DIP-A 002118 , SAMC GoogleMaps

Distribution, biodiversity hotspots, phenology, and biology.

Known from localities in the Northern Cape, Western Cape, and the Eastern Cape of South Africa (Fig. 9 View Figure 9 ). This species is widely distributed and known from 216 specimens, collected in 61 collecting events spread between 1874–2019. The species is known to occur in the Succulent Karoo, Cape Floristic Region, and the Maputaland-Pondoland-Albany biodiversity hotspots. Adult flies are active from August - January (Table 2 View Table 2 ). Adult flies have been observed visiting flowers (https://www.inaturalist.org/observations/64518819). Larvae are unknown but presumed parasitoids as other Bombyliidae , hosts are unknown ( Yeates and Greathead 1997).

Remarks.

Variation in the wing venation connecting R 1 and R 2 + 3 has been observed in a few specimens of this species. Some specimens had 1–3 connecting crossveins (Fig. 10 A View Figure 10 ), instead of the typical 2 crossveins. Coloration is also variable within this species; darker brown specimens and lighter specimens with sandy yellow coloration and more transparent wings, instead of the normal brown have been observed, Hesse (1956) also made note of this variation.