var. griffithii

a. var. griffithii

Lasianthus griffithii Wight (1846) View in CoL 505; Hook.f. (1880) 179; King & Gamble (1904) 111; Merr.(1921) 577; Ridl. (1923a) 152; K.M. Wong (1989) 371. — Type: Kew Dist. 2935 ( Griffith s.n.) (holo K), Peninsular Malaysia.

Lasianthus spathulatus Miq. (1869) View in CoL 246, syn. nov. — Type: Vriese & Teysmann s.n. (holo L, barcode L0057497 ), Borneo.

Lasianthus polycarpus Miq. (1869) 247; Merr. (1921) 579; (1929a) 292; (1937) 298, syn. nov. — Type: Vriese & Teysmann s.n. (holo L, barcode L0057500 ), Borneo.

Lasianthus griffithii Wight var. latibracteata King & Gamble (1904) 112. — Syntypes: Ridley 10196 (n.v.), Selangor; Ridley 9528 (n.v.), Perak; Scortechini 6172 (iso K).

Treelets, up to 5 m high; branchlets terete, robust, c. 3–4 mm diam, glabrous to more or less stiffly hirsute,dark-brown when dry. Leaves: blades oblong-lanceolate, obovate to obovate-lanceolate, 18–25 by 4–9 cm, coriaceous, glabrous to sparsely hirsute above, glabrous to densely hirsute beneath, apex cuspidate to caudate-cuspidate, margin often reflexed, base cuneate to acuminate, midrib and nerves flat to slightly prominent above, prominent beneath, nerves 12– 20 pairs, usually ascending at an angle of over 50°, curved to the margin, nervules obscure to prominent above, prominent beneath, reticulate; petioles 1–2 cm long, glabrous to hirsute. Stipules big, partly covering the cymes, ovate to triangular-ovate or broadly triangular, 1–2 by 1–2 cm, coriaceous with membranaceous margin, usually glabrous. Inflorescences sessile; bracts numerous, linear, hirsute, forming a dense head. Flowers sessile, sealed (hidden) by bracts; calyx tube obconic, c. 2 mm long, glabrous, calyx limb funnel-shaped, 3–4 mm long, glabrous outside, hirsute inside, calyx lobes usually 6, ovate-triangular, c. 2 mm long, glabrous outside, densely hirsute inside, or hirsute both sides; corolla short, c. 4–5 mm long, glabrous outside, hairy inside. Drupes turbinate-ellipsoid, c. 1 cm long, the base narrowed, the apex crowned by a cupular calyx limb, glabrous except the calyx limb; pyrenes usually 6.

Distribution — Malaysia (Peninsular, Borneo), Indonesia (Sumatra).

Notes — Lasianthus griffithii has large obovate-elliptic leaves (glabrous to hairy), large broad-triangular stipules, flowers densely covered by numerous filiform to lanceolate bracts with dense hairs outside, calyx with 5 –7 triangular hairy lobes, drupes teethed. Lasianthus polycarpus is just the same species as L. griffithii . Lasianthus spathulatus differs from L. griffithii by having only relatively narrow, so-called spathulate leaves, which is not enough to separate it from the latter. Wong (1989) sank L. griffithii var. latibracteata .

Lasianthus griffithii is quite variable in leaf sizes and indumentum. It has leaves from relatively narrow spathulate to obovate, from 15 cm long to 35 cm long, most are oblong-obovate and 18– 25 cm long. It is glabrous on branches, leaves and stipules to tomentose or hirsute hairs on leaves beneath, branches and leaves beneath or whole plant.