Lasianthus longifolius subsp. laevis, 2012

b. subsp. laevis H.Zhu View in CoL , subsp. nov. — Fig. 22 View Fig

A typo foliis glabris nitidis, nervulis subtus et supra prominentibus, corollis usque ad 2.5 cm longis,drupis limbis calycis dilatatis cupulatis differt. — Ty- pus: Hansen 295 (holo L), Borneo , Sarawak, Gunong Mulu, 200 m alt., in kerangas forest, 7 Feb. 1978 .

Treelets, branchlets robust, often compressed, glabrous. Leaves: blades obovate-oblong to lanceolate-oblong, 20 –33 by 6 –11 cm, cuspidate to cuspidate-acute at apex, broadly cuneate at base, coriaceous, glabrous and nitid, with the secondary nerves from the costa between nerves, smooth leaf beneath, nerves 7–12 pairs, nervules prominent on both surfaces. Cymes sessile; bracts inconspicuous or fewer, small. Calyx tube obconical, glabrous, limb cupular; corolla tubular, up to 2.5 cm long. Drupes 5– 6 mm diam, crowned by an enlarged cupular calyx limb, ridged; pyrenes 6, ridged on the abaxial face.

Distribution — Borneo ( Sarawak) and Brunei.

Habitat & Ecology — In lowland primary forest below 500 m altitude. Flowering during October to February; fruiting in May.

Additional examined specimens. Sarawak, Ulu Medamit, R. George et al. S.43027 (L); Sarawak, G. Mulu, Paul Chai S. 39588; Sarawak, the 4th Division, Othman et al. S.49087 (K). Brunei, Belait, J. Dransfield 6854 (L); Brunei, Bukit Teraja, M.J.E. Coode 6896 (L), D.A. Simpson 2076.

Notes — This subspecies differs from the typical subspecies by having glabrous and nitid leaves with the secondary nerves from the costa between nerves, smooth leaf beneath, nervules prominent on both surfaces, corolla up to 2.5 cm long, crowned by an enlarged cupular calyx limb. While the typical subspecies has leaves beneath with rigid minute hairs on venation and scabrous (asperate) when touching, nervules conspicuously prominent beneath, corolla less than 1 cm long, drupes crowned by a short cupular calyx limb (not enlarged).

This subspecies was treated as the typical subspecies. However , the specimens from Borneo are different from the ones from Peninsular Malaysia and Sumatra. The specimens from Peninsular Malaysia and Sumatra agree well with the type, while the ones from Borneo are different as discussed above. It is better to recognise the specimens from Borneo as a new subspecies .

From specimens annotated labels, this taxa has calyx white, petals reflexed, with long violet hairs on inner surface.