Cyrteumenes deplanatus, Selis, 2025

Cyrteumenes deplanatus sp. nov.

( Figs 18 View FIGURE 18 , 63D View FIGURE 63 )

Diagnosis. Cyrteumenes deplanatus sp. nov. is immediately distinguished from all other species of Cyrteumenes by the elongate and dorsoventrally depressed mesosoma, with mesoscutum, scutellum and metanotum forming an evenly flattened surface ( Fig. 18C View FIGURE 18 ).

Type material examined. HOLOTYPE: ♀ labelled “ CASENT / 3003723 // MADAGASCAR Fianarantsoa / Prov. Parc Nacional Ranomafana / Bell Vue Trail , tropical forest / 21°15.6'S 47°25.6'E 1000 m / M.E. Irwin and E.I. Schlinger / MEI.99-MA-7 21-XII-1999 // Cyrteumenes deplanatus / HOLOTYPUS ♀ / Det. Marco Selis 2025 [red label]” ( CAS, code CASTYPE21743) GoogleMaps . PARATYPES: Fianarantsoa, PN Ranomafana, Belle Vue at Talatakely, 21.2665S 47.420167E, 1020 m, 28.IV–5. V GoogleMaps .2002, leg. R. Harin’Hala, 1♀ ( CAS, code CASENT2046434 ) .

Description. Female holotype ( Fig. 18B View FIGURE 18 ). Body length 9.0 mm; fore wing length 7.0 mm.

Head as wide as high. Clypeus in frontal view as wide as long, free apical part much shorter than interocular part, apical margin very narrowly emarginate, 0.1× as wide as clypeus, small apical teeth acute and shortly carinate, lateral margins of free apical part sinuate with short lamellate expansion in basal third; clypeus in lateral view very weakly convex, more markedly in basal half. Vertex 1.75× as long as ocellocular distance, posteriorly sloping and very weakly convex in lateral view. Gena 0.9× as wide as eye at bottom of ocular sinus, gena in dorsal view weakly convex and posteriorly converging; occipital carina complete and sharp, finer on vertex and shortly lamellate on gena, strongly bent in lower half. F1 1.4× as long as wide and 1.45× as long as F2, F2 subquadrate and F3–F9 transverse, becoming progressively wider. Mesosoma 1.85× as long as wide, elongate and slightly fusiform in dorsal view, markedly depressed dorsoventrally in lateral view, with mesoscutum, scutellum and metanotum forming an evenly flattened surface smoothly passing into propodeum. Pronotum with straight and anteriorly converging sides, humeri bluntly angled in dorsal view and right-angled in anterior view; pronotal carina complete, sharply lamellate and hyaline on dorsal face, lower and blunter on lateral faces; pretegular carina absent; dorsal and lateral faces of pronotum forming an evenly curved surface, shallowly furrowed along pronotal carina on lateral faces. Mesoscutum longer than wide, with very shallow and wide notauli in front of scutellum. Scutellum subquadrate, slightly wider than long and entirely flattened, anterior margin with a small and shallow median pit; axillary fossa small and rounded, slightly smaller than one ocellus. Metanotum short and flattened, smoothly passing into scutellum and propodeum, posterior margin evenly curved. Tegula campanulate and about as long as wide, with somewhat medially bent outer margin, posterior lobe short and wide with pointed apex equaling parategula; parategula somewhat scythe-shaped, flattened laterolaterally and with sharply pointed apex. Mesepisternum elongate and very low, strongly oblique in lateral view and smoothly passing into wide and flattened mesosternum; epicnemial carina completely absent. Propodeum elongate and with an oblique surface behind metanotum, surface longer than metanotum but not sharply separated from posterior depression of propodeum, medially furrowed; dorsal faces of propodeum evenly and weakly convex, meeting in the middle to form dorsal surface; lateral faces of propodeum flattened and not clearly separated from dorsal faces; posterior face of propodeum forming a circular and deep depression, with a strong longitudinal carina at its bottom; submarginal carina of propodeum strongly developed in a wide and subtriangular lamellar lobe, apically rounded. T1 elongate pyriform, 2.35× as long as apically wide, petiolate and subquadrate in section in basal two-fifths, sub-triangularly expanded and dorsally convex in posterior three-fifths, basal stalk basally tuberculate and then longitudinally carinate in the middle; T 1 in lateral view abruptly raising at extreme base, flattened on basal stalk, convex and becoming progressively higher posteriorly on expanded part, then preapically abruptly furrowed and thickened apically. T2 longer than wide and bell-shaped, constricted basally and much wider apically than basally; apical margin of T2 with a long brownish lamella not markedly separated from rest of surface except for a series of shallow oblique punctures; T 2 in lateral view basally convex and then flattened to apex. S1 elongate bell-shaped, narrow and parallel-sided on basal stalk and then sub-triangularly expanded in posterior half; narrow basal part longitudinally carinate medially, expanded part evenly flattened. S 2 in lateral view obliquely sloping basally and then flattened, the two surfaces separated by an extremely shallow fold of the surface visible only under incident light; apical margin of S2 lamellate similar to T2, but lamella shorter and somewhat more markedly separated from rest of surface.

Frons with deep punctures separated by flattened interspaces ranging from half to one puncture diameter, punctures becoming denser near ocellar triangle; vertex with punctures similar to frons but larger; gena with sparser punctures than frons; clypeus with well impressed circular punctures, denser and deeper basally and much sparser and shallower on disc and apex. Mesosoma with larger and generally sparser punctures; anterior face of pronotum smooth except for some fine punctures near ventral margin; lateral faces of pronotum very sparsely punctate on ventral end; mesoscutum with very shiny and sparsely micropunctate interspaces, becoming wider from anterior to posterior end; scutellum similar to posterior third of mesoscutum; tegula smooth and shiny with very sparse micropunctures; mesepisternum with interspaces equal to puncture diameter on upper end and becoming much wider ventrally; dorsal faces of propodeum with deep punctures separated by very wide shiny interspaces; metaepisternum and lateral faces of propodeum with smaller and sparse punctures, interspaces very finely and not clearly microsculpted, with a silky shine; posterior depression of propodeum with small deep punctures arranged in irregular semicircular series. T1 with some scattered oblique and flat-bottomed punctures on basal stalk, apical expanded part with small deep punctures becoming progressively denser apically but always separated by interspaces as wide as several puncture diameters, thickened apical margin impunctate; T2 basally punctate similar to expanded part of T1, punctures becoming finer and somewhat denser on disc and apex, preapical area of T2 impunctate except for series of shallow punctures bordering the apical lamella; T3–T6 finely micropunctate, micropunctures becoming much finer and sparser from T3 to T6; S1 shiny, basal stalk with some punctures bordering the median carina, apical expanded part with few scattered fine punctures; S2 with larger and much sparser punctures than T2; S3–S6 similar to respective tergites but micropunctures much finer and sparser. Head and mesosoma with long pale-brownish erect setae and short pubescence, setae longer on frons, pronotum, scutellum and propodeum, setae on mesepisternum bent at apex, pubescence denser and silvery on clypeus, ocular sinus and sides of mesosoma; metasoma with very short, dust-like pubescence, erect setae on basal stalk of T1, apex of T5–T6 and S1–S6, setae on basal stalk of T1 and S1 much longer and finer.

Black; following parts light to dark red: mandible, large sub-pentagonal spot on clypeus, apex of scape, wide longitudinal bands along eye margins and in middle on frons, most of pronotum with darkened dorsolateral surfaces, scutellum, tegula, most of mesepisternum, expanded part of T1 and S1, irregular margins of T2, most of S2, irregular apical marking on fore femur, tibiae and tarsi; following parts ivory-yellow: clypeus except median red spot, lower face of scape, interantennal space, narrow anterior band on pronotum becoming slightly wider on humeri, narrow apical bands on T1, T2 and S2; T3–T6 and S3–S6 orange. Wings fusco-hyaline with weak brownish tinge.

Male. Unknown.

Variability. The paratype shows a darker pattern, with much reduced red markings on mesosoma and metasoma.

Distribution. Madagascar: Fianarantsoa ( Fig. 63D View FIGURE 63 ).

Etymology. The species epithet is the Latin word deplanatus (= flattened, made even), in reference to the evenly flattened dorsal surface of the mesosoma.