Paramblynotus yangambicolus (Benoit, 1956)
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publication ID |
https://dx.doi.org/10.3897/jhr.31.4072 |
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publication LSID |
lsid:zoobank.org:pub:DFD1344D-FCA6-42CD-BD68-4FDF2E73F9AC |
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persistent identifier |
https://treatment.plazi.org/id/1C26338B-EFD9-3487-548B-701D101D7E63 |
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treatment provided by |
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scientific name |
Paramblynotus yangambicolus |
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species-group |
Paramblynotus yangambicolus species-group
Remarks.
Historically this taxon has been recognized variously at generic ( Decellea Benoit, 1956) or subgeneric level ( Liu et al. 2007). Decellea was synonymized with Paramblynotus by Weld (1962); followed by re-establishment of generic status by Ronquist (1995); and subsequent re-synonymisation based on phylogenetic analyses, which showed this species-group to be deeply nested within the Paramblynotus clade ( Liu et al. 2007).
Previously the species-group was only known from the African mainland with three described species ( Paramblynotus mixtus Liu et al., Paramblynotus alveolatus Liu et al. and Paramblynotus yangambicolus Benoit). We describe two further species from Madagascar ( Paramblynotus behara sp. n. and Paramblynotus zohy sp. n.).
Diagnosis.
This species-group is characterized in females by excavations (spiracular peritremata) on the terminal portion of T8 associated with the spiracle ( Figs 7B View Figure 7 ; 9 D View Figure 9 ; 12A-B View Figure 12 ), not referred to by Liu et al. (2007). A distinct pronotal crest is present, medially forming a conspicuous, slightly backward pointing, ridge-like tooth ( Figs 4C View Figure 4 ; 6C View Figure 6 ; 8C-D View Figure 8 ; 10C View Figure 10 ). The mesoscutum has rough discontinuous transverse costae produced into irregularly raised and slightly backward pointing teeth ( Figs 4C-D View Figure 4 ; 6C-D View Figure 6 ; 8E-F View Figure 8 ; 10C-D View Figure 10 ). The speculum is longitudinally costate ( Fig. 11D View Figure 11 , and the median propodeal area is not delimited by lateral propodeal carinae.
Key to species of the Paramblynotus yangambicolus species-group (modified from Liu et al., 2007)
| 1 | Forewing with RS+M vein arising at or very close to base of basal vein ( Figs 5B View Figure 5 ; 12C View Figure 12 ) (Madagascar) | 2 |
| - | Forewing with RS+M vein arising at basal third or mid-way up basal vein ( Figs 9B; 9E View Figure 9 ) (Africa) | 3 |
| 2 | Head and mesosoma black ( Figs 4C-D View Figure 4 ). Abdominal tergite 2 (petiole) dorsally sparsely variolate with confused, weak, longitudinal carinulation ( Fig. 4F View Figure 4 ); laterally with strong, widely spaced longitudinal carinae ( Fig. 4E View Figure 4 ). RS+M, basal vein and M+Cu1 nebulous ( Fig. 5B View Figure 5 ). RS+M arising from base of basal vein, this juncture represented by a pigmented spot ( Fig. 5B View Figure 5 ). (Female unknown) | Paramblynotus behara sp. n. |
| - | Head pronotum, mesoscutum and scutellum reddish-brown ( Figs 10 C-D View Figure 10 ). Abdominal tergite 2 (petiole) densely longitudinally carinate ( Fig. 11E View Figure 11 ). RS+M, basal vein and M+Cu1 distinct ( Fig. 12C View Figure 12 ). RS+M arising slightly above base of basal vein ( Fig. 12C View Figure 12 ). (Female T8 with two excavations, a smaller circular one surrounding the spiracle and a second larger oval one adjacent to the first and extending to posterior margin) ( Figs 12A-B View Figure 12 ) | Paramblynotus zohy sp. n. |
| 3 | Body length about 4 mm. Body colour entirely dark ( Fig. 6A View Figure 6 ). Eyes prominent, protruding distinctly beyond genae ( Fig. 6E View Figure 6 ). Median frontal carina almost reaching clypeus. Antennal scrobes rugose. Speculum distinctly longitudinally carinate ( Fig. 6C View Figure 6 ). Upper mesopleuron foveate-reticulate. Mesoscutellum sloped posteriorly ( Fig. 6C View Figure 6 ). Wings entirely transparent ( Fig. 6A View Figure 6 ). Metasoma somewhat, but not strongly, compressed laterally. T6 almost about the same size as the three preceding tergites ( Fig. 6F View Figure 6 ). Sterna 4-6 entirely covered by sternum 3 | Paramblynotus mixtus |
| - | Body length about 6-10 mm. Head and mesosoma dark, metasoma yellow to brown ( Fig. 8A View Figure 8 ). Eyes not protruding distinctly beyond genae ( Fig. 9A View Figure 8 ). Median frontal carina absent in lower face. Antennal scrobes entirely longitudinally carinate. Speculum very finely and superficially carinate ( Fig. 8E View Figure 8 ). Upper mesopleuron entirely longitudinally costate. Mesoscutellum raised posteriorly, forming a flat dorsal surface ( Fig. 8C View Figure 8 ). Forewing evenly ferruginous with darker marginal cell and a dark narrow strip along anterior-interior margin of the first submarginal cell ( Fig. 9B View Figure 9 ). Metasoma strongly compressed laterally ( Fig. 9B View Figure 9 ). T6 distinctly larger than any of the 3 preceding tergites ( Fig. 9C View Figure 9 ). Sterna 4-6 exposed, not covered by sternum 3 | 4 |
| 4 | Face evenly curved in lateral view. Genae ventrally strongly expanded posteriorly. Median mesopleural impression distinct. Lower mesopleuron densely punctate with pubescence. Apical teeth of metatibia rounded apically; first metatarsomere 2.0 × length of remaining 4 metatarsomeres combined | Paramblynotus alveolatus |
| - | Face distinctly raised medially and curved inward ventrally in lateral view ( Fig. 8D View Figure 8 ). Median mesopleural impression usually obscured by extension of longitudinal carinae in upper mesopleuron ( Fig. 8C View Figure 8 ). Lower mesopleuron glabrate and sparsely punctate with sparse pubscence. Apical teeth of metatibia pointed apically ( Fig. 9C View Figure 9 ); first metatarsomere 1.5 × length of remaining 4 metatarsomeres combined | Paramblynotus yangambicolus |
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