Bothynus fabius (Faimaire, 1878)

Bothynus fabius (Faimaire, 1878) View in CoL , here revalidated

( Figures 2 View Figure 2 (j), 4(e), 5(e,g), 11(l), 12(f), 13q, 14q, 16(f), 18(e), 19(b,d), 21(d), 24)

Scaptophilus fabius View in CoL . Fairmaire 1878: 264 (original combination)

Scaptophilus striatellus Fairmaire 1878: 264 View in CoL (original combination)

Scaptophilus quadratus Fairmaire 1878: 265 View in CoL (original combination)

Bothynus fabius View in CoL . Arrow 1937b: 40 (catalogue, as new combination); Blackwelder 1944: 255 (checklist, distributional information); Endrödi 1969: 108 (remarks), 112 (lectotype designation, synonymy proposition)

Bothynus striatellus View in CoL (here synonymised). Arrow 1937b: 41 (catalogue, as new combination); Blackwelder 1944: 255 (checklist, distributional information); Endrödi 1969: 108 (remarks), 112 (identification key), 116 (illustration of the aedeagus [figs. 158–159]), 131–132 (redescription, lectotype designation, distribution); Endrödi 1985: 274 (illustration of the aedeagus [figs. 1112–1123], identification key, distribution); Krajcik 2005: 39 (checklist); Abadie et al. 2008, pl. 14 ( Figure 7 View Figure 7 , distribution); López-García et al. 2016: 497 (checklist, distribution).

Bothynus quadratus View in CoL . Arrow 1937b: 41 (as new combination); Blackwelder 1944: 255 (checklist, distributional information); Endrödi 1969: 108 (synonymy proposition).

Diagnosis

Bothynus fabius resembles B. deiphobus in the inner protarsal claw of male simple, equal to outer claw, without incision or ventromedial process, glabrous and deep-striated elytra, and parameres without ventrobasal carina. However, B. fabius is distinguished from B. deiphobus by the following characters: Hemisternite separated by a partial or complete longitudinal midline, never entire fused ( Figure 12 View Figure 12 (f)); metatarsomere 1 with short outer apex in both sexes ( Figure 18 View Figure 18 (e)); anterior half of tergite 7 with 1 band of well-marked stridulatory striae in both sexes ( Figure 19 View Figure 19 (b)); females bearing a cavity on each side of tergite 8 ( Figure 19 View Figure 19 (d)).

Type material

Bothynus striatellus . Lectotype female, labelled: (a) ‘ Bothynus striatellus /Entrerios Fairm. Type’. [handwritten]; (b) ‘Museum Paris/ex Coll./R.Oberthur’ [light green label, bordered in black]; (c) ‘ Lectotypus /Both (Scaptoph)/striatellus/Fairm/Endrödi’ [label bordered in red, partly handwritten]; (d) ‘MNHN/EC7110’ ( MNHN) ( Figure 2 View Figure 2 (j)). Paralectotype male, labelled: (a) ‘EntreRios/Dohrm’ [handwritten]; (b) ‘Scapth./striatellus/Fairm./Plata’ [handwritten]; (c) ‘MUSEUM PARIS/Collection Léon Fairmaire/1906’; (d) ‘ Paratypus /Both. (Scaptoph)/striatellus/Fairm’. [label bordered in red, partly handwritten]; (e) ‘PARALECTOTYPE’ [red label”; (c) ‘MNHN/EC7111’ ( MNHN). Bothynus fabius . Lectotype male, labelled: (a) ‘ Scaptophilus /fabius/Farm/am.mer’. [handwritten]; (b) ‘MUSEUM PARIS/ Collection Léon Fairmaire/1906’ [light green label]; (c) ‘TYPE’ [red label]; (d) ‘ Lectotypus / Bothynus (Scapt.) / fabius Fairm /Endrödi’ [label bordered in red, partly handwritten]; (e) ‘penis/default!!’ [handwritten]; (f) ‘MNHN/EC7112’ ( MNHN).