Pithocerithium longiuscatum ( Sacco, 1895 )

Pithocerithium longiuscatum ( Sacco, 1895)

Figs 27H, 33A–E View FIGURE 33

Cerithium doliolum Brocc.— Hörnes 1855: 392 View in CoL (pars), pl. 41, fig. 12 [non Pithocerithium doliolum ( Brocchi, 1814) ]. * [ Cerithium (Pithocerithium) turonicum View in CoL ] var. longiuscata Sacco 1895: 29 . Cerithium exdoliolum Sacco— Friedberg 1914: 265 , pl. 16, fig. 14 [non Pithocerithium exdoliolum ( Sacco, 1895) ]. Cerithium (Pithocerithium) turonicum Mayer— Sieber 1937: 500 View in CoL , pl. 25, fig. E2 [non Pithocerithium turonicum (Mayer,

1878)]. Cerithium (Vulgocerithium) exdoliolum Sacco — Csepreghy-Meznerics 1950: 31, pl. 2, fig. 4 [non Pithocerithium exdoliolum

( Sacco, 1895)]. Cerithium (Thericium) exdoliolum Sacco— Korobkov 1955 : plate captions, pl. 26, fig. 4 [non Pithocerithium exdoliolum ( Sacco,

1895)]. Cerithium (Vulgocerithium) exdoliolum Sacco 1895 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 106, pl. 31, fig. 8

[non Pithocerithium exdoliolum ( Sacco, 1895) ]. Cerithium (Thericium) boettgeri nom. nov. — Švagrovský 1960: 77, pl. 7, figs 11–12. Cerithium cfr. exdoliolum Sacco— Strausz 1962: 230 , pl. 9, figs 27–28 [non Pithocerithium exdoliolum ( Sacco, 1895) ]. Cerithium cfr. exdoliolum Sacco— Strausz 1966: 552 , pl. 9, figs 27–28 [non Pithocerithium exdoliolum ( Sacco, 1895) ]. Cerithium (Thericium) turonicum turonicum Mayer— Mariani & Papp 1966: 143 , pl. 1, figs 1–2 [non Pithocerithium turonicum

(Mayer, 1878)]. Cerithium exdoliolum Sacco, 1895 — Zelinskaya et al. 1968: 154, pl. 38, fig. 8 [non Pithocerithium exdoliolum ( Sacco, 1895) ]. Cerithium vulgatum europaeum Mayer, 1878 — Nicorici & Sagatovici 1973: 165, pl. 24, fig. 2 [non Thericium europaeum

(Mayer, 1878)]. Cerithium (Thericium) aff. zelebori Hörnes, 1856 — Bałuk 1975: 147, pl. 17, figs 4–5 [= non Hörnes, 1855]. Cerithium (Thericium) exdoliolum ( Sacco, 1895) — Švagrovský 1982: 20, pl. 9, fig. 1 [non Pithocerithium exdoliolum ( Sacco,

1895)]. Cerithium (Thericium) turonicum Mayer— Kókay 1996: 456 , pl. 3, fig. 2 [non Pithocerithium turonicum (Mayer, 1878) ]. Thericium longiuscatum ( Sacco, 1895) — Landau et al. 2013: 42, pl. 3, fig. 9.

non Cerithium doliolum Brocc. var. longiuscata Sacc.— Bauer 1900: 30 , pl. 1, fig. 10. [= Thericium sp. ].

Type material. Holotype, NHMW 1999 View Materials z0098/0008, SL: 24.9 mm, MD: 9.9 mm, Steinebrunn ( Austria), illustrated in Hörnes (1855: pl. 41, figs 12a–b), Figs 33A View FIGURE 33 1 –A View FIGURE 1 2 View FIGURE 2 .

Illustrated material. NHMW 1870/0037/0037, SL: 25.7 mm, MD: 8.7 mm, Ritzing ( Austria), Figs 27H, 33B View FIGURE 33 1 – B View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0024/0001, SL: 27.1 mm, MD: 9.3 mm, Ritzing ( Austria), Figs 33C View FIGURE 33 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0024/0002, SL: 24.9 mm, MD: 8.4 mm, Ritzing ( Austria), Figs 33D View FIGURE 33 1 –D View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0024/0003, SL: 24.8 mm, MD: 9.6 mm, Ritzing ( Austria), Figs 33E View FIGURE 33 1 –E View FIGURE 1 2 View FIGURE 2 .

Additional material. 6 spec., NHMW 1884 View Materials /0018/1612, Guntersdorf ( Austria) ; 13 spec., NHMW 2023 View Materials /0024/0004, Ritzing ( Austria) ; 1 spec., NHMW 1999 View Materials z0004/0018, Ritzing ( Austria) , illustrated in Sieber (1937: pl. 25, fig. E2); 14 spec., NHMW 1930 View Materials /0006/0046, Ritzing ( Austria) ; 2 spec., NHMW 1874 View Materials /0033/0016, Lăpugiu de Sus ( Romania) .

Revised description. Medium sized, elongate pupoid shell of up to ten teleoconch whorls, attaining ~ 24– 27 mm in height; apical angle 40–45°. Protoconch unknown. Early teleoconch whorls low, weakly convex with faint mid-whorl angulation and wide spaced, drop-shaped axial ribs, crossed by delicate spiral cords. Later whorls subcylindrical to weakly convex. Axial ribs disintegrating into weak subsutural spiral row of rounded tubercles and spiral cord of horizontally elongated tubercles placed just above suture, separated by weakly concave subsutural ramp. Additional spiral cord of weaker tubercles appears around eighth teleoconch whorl at abapical suture. Single narrow secondary spiral thread intercalated between broad primary spiral cords. Suture deep, narrow, more or less linear. Last whorl high, ovate, attaining ~45% of total height, with prominent varix at about 120° from the edge of the outer lip Sculpture of five tubercular cords, adapical broadest, with 3–4 crowded spiral threads intercalated in interspaces. Base weakly constricted with two weakly beaded spiral cords and several prominent secondary spirals. Aperture ovate, wide. Columella excavated. Columellar callus forming broad, thick rim, sharply delimited from base. Anal canal deeply incised, moderately narrow, with prominent parietal denticle. Outer lip slightly thickened. Siphonal canal short, wide, deflected to the left.

Synonyms. Cerithium (Thericium) boettgeri Švagrovský, 1960 . Holotype illustrated in Švagrovský 1960: 77, pl. 7, fig. 12, Department of Geology and Paleontology, Faculty of Natural Sciences Comenius University Bratislava. Slanské hory ( Slovakia), Middle Miocene, late Badenian (Serravallian). This species seems to be based on a slightly aberrant specimen of P. longiuscatum with rather pointed and wide spaced tubercles.

Discussion. The name longiuscatum was overlooked so far in the literature because it was introduced by Sacco (1895: 29) somewhat hidden within a discussion on P. turonicum (Mayer, 1878) referring to the specimen illustrated as fig. 12 in Hörnes (1855: pl. 41).

This species is characterized by its slender pupoid outline and regularly beaded sculpture. It is distinguished from P. turonicum (Mayer, 1878) by its larger size and the elongate and slightly cyrtoconoid spire. Morphologically, Pithocerithium longiuscatum ( Sacco, 1895) is closer to P. mediolanionense nom. nov. than to P. turonicum . Pithocerithium mediolanionense differs in its broader ovate outline and differences in sculpture, such as the large subquadratic beads on the subsutural and mid-whorl cords. Pithocerithium longiuscatum might be related to Pithocerithium praedoliolum ( Fischer & Tournouër, 1873) from the Tortonian of Vaucluse ( France), which has a comparably slender outline, but differs in its larger but fewer and wide spaced tubercles and wider interspaces between the primary spiral cords (see Fischer & Tournouër 1873: pl. 18 fig. 1; https://science.mnhn.fr/institution/ mnhn/collection/f/item/r06717).

Paleoenvironment. Shallow marine, inner neritic (own data M.H.).

Distribution. Pithocerithium longiuscatum was widespread in the Badenian of the Central Paratethys Sea and occurred also in the Serravallian of the Proto-Mediterranean Sea.

Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Bałuk 1975); Polish-Ukrainian Fore-Carpathian Basin: Wólka Niedźwiedzka, Zgłobice ( Poland), Zaichyky (Khmel’nytskyi Region), Khorostkiv, Rydomyl’ (Ternopil Region, Ukraine) ( Friedberg 1914, hoc opus); North Alpine Foreland Basin: Guntersdorf( Austria) (hoc opus); Oberpullendorf Basin: Ritzing ( Austria) (hoc opus); Vienna Basin: St. Veit/Berndorf, Steinebrunn ( Austria), Mikulov ( Czechia), Borský Mikuláš ( Slovakia) (hoc opus; Švagrovský 1982); Danube Basin: Modra-Kráľová ( Slovakia) ( Švagrovský 1982); Eastern Slovakian Basin: Slanské hory ( Slovakia) ( Švagrovský 1960); Pannonian Basin: Hidas, Márkháza, Sámsonháza, Várhegy ( Hungary) ( Csepreghy-Meznerics 1950, 1954; Strausz 1966); Făget Basin: Lăpugiu de Sus ( Romania) (hoc opus); Zarand Basin: Minişul de Sus ( Nicorici & Sagatovici 1973); Dacian Basin: Opanets ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Seyithasan ( Turkey) ( Landau et al. 2013).