Pithocerithium mediolanionense, Harzhauser & Guzhov & Landau, 2025

Pithocerithium mediolanionense nom. nov.

Figs 27J, 34A–G View FIGURE 34

Cerithium doliolum Brocc.— Hörnes 1855: 392 View in CoL (pars) [non Pithocerithium doliolum ( Brocchi, 1814) ].

Cerithium (Pithocerithium) doliolum Brocc. var. imperfecta n. v. — Sieber 1937: 499, pl. 25, fig. E3 [non Deshayes, 1833].

C [erithium]. (Th [ericium].) italicum May.— Sieber 1958: 137 [non Pithocerithium italicum (Mayer, 1878) ].

Cerithium (Thericium) turonicum imperfecta Sieber 1937 — Mariani & Papp 1966: 143 [non Deshayes, 1833 = Hemicerithium imperfectum View in CoL ].

Cerithium turonicum (Mayer, 1878) View in CoL —Harzhauser 2002: 72, pl. 2, figs 5–8 [non Pithocerithium turonicum (Mayer, 1878) ].

Clavatula (Clavatula) granulatocincta View in CoL (Münster in Goldfuss, 1840) — Bošnjak et al. 2021: 228, fig. 3A [non Clavatulidae View in CoL ].

Type material. Holotype, NHMW 2023/0025/0001, Niederkreuzstetten ( Austria), SL: 20.5 mm, 8.5 mm, syntype of Cerithium (Pithocerithium) doliolum imperfecta Sieber, 1937 , illustrated in Sieber (1937: pl. 25, fig. E3), Fig. 34C View FIGURE 34 .

Illustrated material. NHMW 1861 View Materials /0050/0149, SL: 26.9 mm, MD: 10.6 mm, Kleinebersdorf ( Austria), Figs 34A View FIGURE 34 1 –A View FIGURE 1 2 View FIGURE 2 . Paratypes : NHMW 2023 View Materials /0026/0001, SL: 27.7 mm, MD: 10.3 mm, Kleinebersdorf ( Austria), Figs 34B View FIGURE 34 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 2023 View Materials /0026/0002, SL: 21.9 mm, MD: 8.2 mm, Kleinebersdorf ( Austria), Figs 34D View FIGURE 34 1 –D View FIGURE 1 2 View FIGURE 2 . NHMW 2023 View Materials /0026/0003, SL: 18.9 mm, MD: 7.8 mm, Kleinebersdorf ( Austria), Fig. 34E–F View FIGURE 34 . NHMW 2023 View Materials /0026/0004, SL: 18.7 mm, MD: 8.9 mm, Kleinebersdorf ( Austria), Fig. 34G View FIGURE 34 ; NHMW 2023 View Materials /0026/0005, SL: 20.7 mm, MD: 8.1 mm, Kleinebersdorf ( Austria), Fig. 27J .

Additional material. 14 spec., NHMW 2023 View Materials /0026/0006, Kleinebersdorf ( Austria) ; 11 spec., NHMW 2023 View Materials /0026/0007, Kleinebersdorf ( Austria) ; 6 spec., NHMW 1861 View Materials /0050/0014, Rückersdorf ( Austria) ; 2 spec., NHMW 1860 View Materials /0050/0064, Neuruppersthal ( Austria) ; 1 spec., NHMW 1859 View Materials /0045/0257, Weinsteig ( Austria) ; 6 spec., NHMW 1861 View Materials /0050/0167, Gebmannsberg ( Austria) ; 5 spec., NHMW 1859 View Materials /0045/0070, Niederkreuzstetten ( Austria) ; all Karpatian (Early Miocene).

Etymology. After the historical village Mediolanion, which was situated in the Korneuburg region in Austria.

Description. Medium sized, stocky, ovate shell of up to nine teleoconch whorls, attaining ~ 18–27 mm in height; apical angle 43–55°. Protoconch unknown. Early teleoconch whorls low with weak mid-whorl angulation and large, knob-shaped axial ribs and delicate spiral threads. Narrow, slightly concave subsutural ramp developing around forth to fifth teleoconch whorl. Axial ribs subobsolete over subsutural ramp. Subsutural spiral cord and weak spiral cord at abapical suture developing on subsequent whorls. Tubercles on central spiral cord become more densely spaced forming close-set beads. Subsequent whorls with three distinct beaded spiral cords separated by narrow interspaces with delicate single secondary spiral thread intercalated. Adapical spiral cord with densely spaced, rectangular, convex, slightly axially elongated beads; central cord with slightly wider spaced, rounded, more prominent, higher tubercles; abapical spiral cord with smaller, horizontally elongated beads. Penultimate whorl with fourth beaded spiral cord appearing at abapical suture. Whorl profile subcylindrical to weakly convex. Suture narrowly incised. Last whorl high, ovate, attaining ~55% of total height. Broad varices on last two whorls. Base weakly constricted, with numerous prominent spiral cords with narrower secondaries intercalated. Aperture only partly preserved. Columella excavated. Columellar callus forming broad, slightly thickened rim, sharply delimited from base. Anal canal distinctly incised, relatively wide, with prominent parietal denticle. Outer lip not preserved. Siphonal canal moderately short, slightly twisted.

Synonyms. Cerithium imperfectum Sieber, 1937 is primary homonym, preoccupied by Deshayes (1833: 365) and therefore, we propose Pithocerithium mediolanionense nom. nov. as replacement name.

Discussion. Pithocerithium mediolanionense nom. nov. is highly reminiscent of P. doliolum ( Brocchi, 1814) from the Pliocene of the Mediterranean Sea concerning the subquadratic beads on the subsutural spiral cord and on the mid-whorl cord. However, they are distinguished by the larger size and the higher and more conical spire of P. doliolum (see Pinna & Spezia 1978: pl. 34, fig. 2; Chirli 2006: pl. 34, figs 9–15). Pithocerithium mediolanionense differs from P. longiuscatum ( Sacco, 1895) in its stout outline and the much broader subsutural beads and larger tubercles on the mid-whorl spiral cord. It differs from Pithocerithium turonicum (Mayer, 1878) in its cyrtoconoid spire, higher last whorl, less constricted base and larger, more close-set beads. A Late Miocene record of this group from the Northeastern Atlantic was described by Pereira Da Costa (1867; 247, pl. 28, fig. 14) from the Tortonian of Cacela ( Portugal) but the poor drawing does not allow an identification.

Paleoenvironment. Coastal marine, lagoonal environments in vicinity of mudflats and Avicennia mangroves (Harzhauser et al. 2002).

Distribution. Mainly known from the Karpatian (Late Burdigalian) of the Central Paratethys Sea and very rare during the Badenian.

Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Gebmannsberg, Helfens, Karnabrunn, Kleinebersdorf, Neuruppersthal, Niederkreuzstetten, Obergänserndorf, Rückersdorf, Stetten, Teiritzberg, Weinsteig ( Austria) (Harzhauser 2002; hoc opus). Badenian (Middle Miocene): Pannonian Basin: Zaprešić Brijeg ( Croatia) ( Bošnjak et al. 2021).