Hemicerithium banaticum ( Boettger, 1902 )

Hemicerithium banaticum ( Boettger, 1902) View in CoL

Figs 65A–C View FIGURE 65

* Cerithium (Conocerithium) banaticum n. sp. — Boettger 1902: 118.

Cerithium (Conocerithium) banaticum Boettger— Zilch 1934: 221 , pl. 8, fig. 33.

Tritonium (?) zboroviense Friedb.— Friedberg 1938a: 113 , text fig. 34.

Cerithium (Conocerithium) banaticum Boettger — Csepreghy-Meznerics 1954: 23, pl. 1, figs 6–7, 13–14.

Conocerithium banaticum Boettger— Strausz 1962: 47 , pl. 10, figs 6–7, text fig. 60.

Conocerithium banaticum Boettger, 1901 — Strausz 1966: 139, pl. 10, figs 6–7.

Cerithium (Conocerithium) banaticum Boettger— Kókay 1966: 45 , pl. 4, figs 19–20.

Cerithium (Conocerithium) banaticum (Boettger) — Csepreghy-Meznerics 1972: 21, pl. 3, fig. 23.

Hemicerithium (Hemicerithium) banaticum (Boettger) View in CoL — Bałuk 1975: 153, pl. 18, figs 4–5.

Conocerithium (Conocerithium) banaticum (Boettger, 1901) — Atanacković 1985: 108, pl. 26, figs 6–7.

Hemicerithium (Hemicerithium) cf. banaticum (Boettger, 1901) View in CoL — Nikolov 1993: 68, pl. 3, figs 1–2.

non Conocerithium banaticum ( Boettger, 1902) — Švagrovský 1981: 133, pl. 41, fig. 9. [= Thericium zeuschneri ( Pusch,

1836)]. non Cerithium (Hemicerithium) banaticum (Boettger, 1901) — Iljina 1993: 82, pl. 10, figs 16–18 [= Hemicerithium caucasicum

sp. nov.].

Type material. Lectotype SMF XII 2427 a, SL: 8.5 mm, MD: 4.1 mm, Senckenberg Forschungsinstitut und Naturmuseum Frankfurt am Main. ( Germany), designated by Zilch (1934: 221, pl. 8, fig. 33), Figs 65A View FIGURE 65 1 –A View FIGURE 1 2 View FIGURE 2 . Coşteiu de Sus ( Romania), Middle Miocene, early/middle Badenian . Paralectotypes SMF XII 2427 b–d, Coşteiu de Sus ( Romania) .

Illustrated material. NHMW 2023/0043/0004, SL: 9.8 mm, MD: 4.9 mm, Coşteiu de Sus ( Romania), Figs 65C View FIGURE 65 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0027/0003, SL: 8.0 mm, MD: 3.9 mm, Baden ( Austria), Figs 65B View FIGURE 65 1 –B View FIGURE 1 2 View FIGURE 2 .

Additional material. 12 spec., NHMW 2023 View Materials /0043/0005, Coşteiu de Sus ( Romania) ; 23 spec., NHMW 2018 View Materials /0248/0557, Coşteiu de Sus ( Romania) ; 12 spec, NHMW 2023 View Materials /0027/0004, Baden ( Austria) ; 2 spec., NHMW 2023 View Materials /0030/0002, Forchtenau ( Austria) ; 2 spec., NHMW 2023 View Materials /0053/0001, Porzteich ( Czechia) ; 3 spec., NHMW 2016 View Materials /0177/0382, Lăpugiu de Sus ( Romania) ; 1 spec., NHMW 2016 View Materials /0177/0395, Lăpugiu de Sus ( Romania) .

Revised description. Small, broadly conical shell of nine teleoconch whorls with prominent, strongly swollen varices, attaining ~ 8–10 mm in height; apical angle ~35°. Protoconch unknown. Early teleoconch whorls convex with delicate spiral sculpture. Later whorls with broad, straight to weakly convex sutural ramp, angled at periphery placed short distance above abapical suture. Sculpture of broad, wide-spaced, irregular axial ribs; spiral sculpture of prominent primary and secondary cords. Periphery in abapical third of whorls, often coinciding with two more prominent spiral cords. Sutural ramp sometimes weakly cancellate. Axial ribs may form broad, low, tubercles at intersections at periphery. Suture deeply incised, undulating. Last whorl convex, inflated, attaining ~45% of total height, with weak peribasal spiral cord. About four varices on last whorl. Base strongly constricted, weakly convex; sculpture of close-set primary and secondary cords covers entire base and fasciole.Aperture wide, ovate. Columellar callus forming narrow rim, weakly delimited from base. Columella straight. Anal canal indistinct. Outer lip thin, smooth. Siphonal canal short, wide, slightly deflected to the left.

Discussion. This species displays some variability concerning slenderness of the late teleoconch whorls and strength of the varices and axial ribs, resulting in specimens with dominant spiral sculpture (e.g., Bałuk 1975, pl. 10, fig. 4). Hemicerithium perrettai Pacaud, 2019 , from the Bartonian of France, is comparable in outline and develops prominent varices, but differs in its reduced axial ribs and weaker spiral sculpture (see Pacaud 2019: pl. 1, fig. 6). Specimens described by Iljina (1993) as this species from the Middle Miocene of the Eastern Paratethys are excluded herein from H. banaticum ; see Hemicerithium caucasicum sp. nov.

Paleoenvironment. Unknown.

Distribution. Only known from the Badenian (Langhian, early Serravallian) of the Central Paratethys Sea.

Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1975); Polish-Ukrainian Fore-Carpathian Basin: Zboriv (Ternopil Region) ( Ukraine) ( Friedberg 1938a); Vienna Basin: Porzteich ( Czechia), Baden ( Austria) (hoc opus); Styrian Basin: Wetzelsdorf ( Austria) ( Strausz 1966); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (hoc opus); Pannonian Basin: Sámsonháza ( Hungary) ( Strausz 1966), Hrvaćani ( Bosnia and Hercegovina) ( Atanacković 1985); Cserhát Hills: Kelet-Cserhát ( Hungary), Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus ( Romania); Dacian Basin: Yasen ( Bulgaria) ( Nikolov 1993).