Hemicerithium evae ( Boettger, 1902 )

Hemicerithium evae ( Boettger, 1902)

Figs 65D–E View FIGURE 65

* Cerithium (Conocerithium) evae sp. nov. — Boettger 1902: 118.

Cerithium (Conocerithium) evae Boettger— Boettger 1906: 139 .

Cerithium (Conocerithium) evae Boettger— Zilch 1934: 221 , pl. 8, fig. 32.

? Tritonium zboroviense Friedb.— Krach & Ksiażkiewicz 1950: 285 .

Conocerithium (Conocerithium) olgae (Boettger, 1901) — Atanacković 1985: 109, pl. 26, figs 8–9 [non Hemicerithium olgae ( Boettger, 1902) ].

Type material. Lectotype SMF XII 2426 a, SL: 12.8 mm, MD: 6.5 mm, Senckenberg Forschungsinstitut und Naturmuseum Frankfurt am Main ( Germany), designated by Zilch (1934: 221), Coşteiu de Sus ( Romania), Middle Miocene, early/middle Badenian, Figs 65D View FIGURE 65 1 –D View FIGURE 1 2 View FIGURE 2 . Paralectotypes SFM XII 2426 b–d, Coşteiu de Sus ( Romania) .

Illustrated material. NHMW 2023/0042/0008, SL: 20.2 mm, MD: 8.6 mm, Lăpugiu de Sus ( Romania), Figs 65E View FIGURE 65 1 –E View FIGURE 1 2 View FIGURE 2 .

Additional material. 6 spec., NHMW 1863 View Materials /0016/0197, Lăpugiu de Sus ( Romania) .

Revised description. Medium sized, broad conical shell of up to 11 teleoconch whorls with weak varices, attaining ~ 13–21 mm in height; apical angle 41–45°. Protoconch unknown. Earliest teleoconch whorls convex with delicate spiral cords. Third teleoconch whorl with mid-whorl angulation and faintly concave sutural ramp; weakly convex below angulation. Whorls covered by fine spiral cords, with close-set pair of more prominent spiral cords at periphery. Axial ribs subobsolete on sutural ramp, forming weak tubercles over peripheral pair of spirals. Suture distinctly incised. Angulation fading on penultimate and last whorls, which become more convex. Surface of last whorls covered by narrow primary and secondary spiral cords. Last whorl convex, attaining ~45% of total height. Varices weak (three on last whorl). Base convex, strongly constricted, delimited by only slightly strengthened peribasal cord. Aperture wide, ovate. Columella moderately excavated. Columellar callus forming narrow rim, poorly delimited from base. Anal canal indistinct. Outer lip smooth. Siphonal canal short, wide, slightly bent to the left.

Synonyms. Tritonium (?) zboroviense Friedberg, 1938 is based on at least two syntypes from the Badenian of Zboriv ( Ukraine). Friedberg (1938, fig. 34) illustrated two specimens of which the right one corresponds fully with H. evae whereas the left one is rather slender and lacks an angulation of the whorls. Herein we designate the right specimen as lectotype of Tritonium (?) zboroviense and consider it to be a subjective junior synonym of H. evae .

Discussion. This species is variable concerning slenderness and size. The holotype represents a relatively small and broad specimen. Bałuk (1975) considered Hemicerithium evae ( Boettger, 1902) to be a subjective synonym of H. banaticum ( Boettger, 1902) , with which it co-occurs. However, they are distinguished by the much larger size of H. evae , the wider apical angle, the much weaker varices, and the angulated periphery of H. evae . Hemicerithium evae is slightly reminiscent of Hemicerithium adela ( Dall, 1915) , from the Early Miocene of Florida ( USA), but differs in its characteristic pair of spiral cords at the periphery (see Hoerle 1972: pl. 2, fig. 2).

Paleoenvironment. The occurrences in the Baden Formation (Baden) suggest middle to outer neritic environments with up to 210 m water depth ( Kranner et al. 2021).

Distribution. Only known from the Badenian (Langhian, early Serravallian) of the Central Paratethys Sea.

Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Porzteich ( Czechia), Baden ( Austria) (hoc opus); Pannonian Basin: Hrvaćani ( Bosnia and Hercegovina) ( Atanacković 1985); Făget Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Zilch 1934: hoc opus).