Parasyntexis khasurtensis Kopylov, 2019

Li, Yi, Rasnitsyn, Alexandr P., Shih, Chungkun, Ren, Dong & Wang, Mei, 2024, New fossil Anaxyelidae (Insecta: Hymenoptera: Siricoidea) from the Lower Cretaceous Yixian Formation of northeastern China, Zootaxa 5562 (1), pp. 107-121 : 114-116

publication ID

https://doi.org/10.11646/zootaxa.5562.1.13

publication LSID

lsid:zoobank.org:pub:F4559678-00E0-4150-82D1-CE9F6AE4C147

DOI

https://doi.org/10.5281/zenodo.14596590

persistent identifier

https://treatment.plazi.org/id/257FAE5A-7438-FFEB-FF51-74FCFE81A61A

treatment provided by

Plazi

scientific name

Parasyntexis khasurtensis Kopylov, 2019
status

 

Parasyntexis khasurtensis Kopylov, 2019

Material. CNU-HYM-LB2024503(P/C).

Locality and horizon. Huangbanjigou, Chaomidian Village, Shangyuan Township, Beipiao City, Liaoning Province, China; Lower Cretaceous (Barremiane–Aptian), Yixian Formation.

Description. Female. Head and thorax predominantly dark, slightly darker than abdomen; ovipositor slightly darker; legs lightly colored; forewing with pterostigma completely sclerotized; cell 1r darkened and H-shaped pattern around veins 1-Cu, 2-Cu, 3-Cu, 1m-cu and 1cu-a dark ( Fig. 3 View FIGURE 3 ).

Head: Head moderately large, nearly oval, narrower than thorax. Compound eye large and ovate ( Fig. 3A, C View FIGURE 3 ). Antenna thick, strongly tapering to sharp apex, apparently short, with fifteen flagellomeres visible; scape slightly expanded ( Fig. 3B View FIGURE 3 ); scape 0.68 mm long, maximum width 0.45 mm; pedicel 0.82 mm long, maximum width 0.19 mm; flagellomere I shorter than scape, 0.21 mm wide, 0.91 mm long; flagellomere II 0.17 mm wide, 0.65 mm long; the visible parts of flagellomeres gradually shortened and their length-width ratio (excluding the first and second flagellomeres) from 3.2 times to one times toward the apex, except for the last flagellomere slender and twice as long as wide.

Thorax: Pronotum trapezoid ( Fig. 3A View FIGURE 3 ). Mesoscutum large, with anterior margin nearly straight, median longitudinal sulcus and notauli strongly impressed; mesoscutellum tapering to acute apex; ratio of lengths of prescutum, median longitudinal sulcus between notauli and mesoscuto-mesoscutellar sulcus, and mesoscutellum 3.7/4.5/4.1 ( Fig. 3B View FIGURE 3 ), mesoscutellum triangular, tapering to acutely sharp apex ( Fig. 3F View FIGURE 3 ).

Leg: Leg with pro- and metafemur shorter than tibia; profemur length 2.01 mm, metafemur length 1.78 mm, thick subapically, narrowed apically. Tarsi long and thin ( Fig. 3B, C View FIGURE 3 ).

Wing: Forewing three times as long as wide ( Fig. 3G View FIGURE 3 ). Vein C and R thick, costal area wide medially. 1r-rs 1.5 times as long as 2r-rs ( Fig. 3H View FIGURE 3 ), 2r-rs issuing from pterostigma at its one-third of apex, as long as pterostigma wide; 1-Rs long, aligned with Rs+M, about 3.2 times as long as 1-M; 2-Rs+M short, 0.2 times as long as 1-Rs+M; 2r-rs slightly proclival; 2+3-M 1.8 times as long as 4-M; 4-M slightly longer than 3r-m and 2m-cu. Cell 1mcu pentagonal, about 1.7 times as long as wide; cell 2mcu about 0.2 times as long as wide; 2-Cu 0.8 times as long as 1-Cu; 1cu-a straight, shorter than 1-Cu; 2A+3A with long subbasal loop; cell 2+3rm long and pentagonal, about 3.2 times as long as wide; cell 1+2r 2.4 times as long as wide; cell 3r about 3.3 times as long as wide; cell 3r as long as 1+2r.

Abdomen: Abdomen slightly distorted, wider than the mesothorax, parallel-sided before ovipositor base. First abdominal tergite split medially ( Fig. 3A, C View FIGURE 3 ). Ovipositor with first valvifer small, subhemicircular as visible; second valvifer large, with thickening medially and distally, narrowing up to single thickening before meeting with valvula 3 (saw sheath), 1.2 times as long as valvula 3, valvula 2 (dorsal stylet) visible only basally at its thickened base ( Fig. 3D, E View FIGURE 3 ); ovipositor straight, sword-shaped ( Fig. 3A, D View FIGURE 3 ); ovipositor full length equal to 1.1 times forewing length; ovipositor elongated and narrowly rounded apically, prominent significantly behind abdomen, valvula 3 0.4 times as long as forewing length.

Measurements (in mm). Body excluding ovipositor about 14.81 in length; head about 2.42 in length, 2.51 in width; thorax about 4.02 in length; abdomen about 8.28 in length; forewing (incompletely preserved) at least 9.72 in length, maximum width 3.61; ovipositor about 4.41 in length.

Remarks. The new specimen can be assigned to the genus Parasyntexis based on the forewing characters: costal area wide; pterostigma as wide as vein 2r-rs long; 1-Rs as long as pterostigma; both 1r-rs and 2r-rs developed, 1r-rs 1.3 times as long as 2r-rs; cell 2r wider than pterostigma; Rs+M furcating after 1m-cu, 2Rs+M present, cell 1mcu pentagonal. Kopylov (2019) described the type species P. khasurtensis based on one specimen; however, the antennae, thorax and ovipositor were not described due to the lack of preservation of these structures. Our well-preserved new specimen (CNU-HYM-LB2024503(P/C)) reported herein can supplement some new body characters for further study.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

InfraClass

Lower

Order

Hymenoptera

SubOrder

Symphyta

SuperFamily

Siricoidea

Family

Anaxyelidae

SubFamily

Syntexinae

Genus

Parasyntexis

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