Dentilla dichroa ( Sichel et Radoszkowski, 1869 )

3. Dentilla dichroa ( Sichel et Radoszkowski, 1869)

( Figs 24–28 View FIGURES 24–32 , 33–35 View FIGURES 33–38 )

Mutilla bicolor Olivier, 1811: 52 , 57, ♂, nom. praeocc., non Pallas, 1771, holotype, ♂: "dans le petit désert de l'Arabie, près de l'Euphrate" [ Iraq] [MNHN].

Mutilla dichroa Sichel et Radoszkowski, 1869: 159 ; 1870: 300, ♂ (replacement name for Mutilla bicolor Olivier, 1811 ; André 1898: 6, ♂.

Mutilla (Ephutomma) dichroa : André 1901a: 150, ♂ (Turkestan, Région transcaspienne, Arabie, Sinai).

Ephutomma dichroa : Skorikov 1935: 324, tab. 5, fig. 1, ♂.

Smicromyrme (Dentilla) dichroa : Lelej & Kabakov 1980: 195, ♂.

Dentilla dichroa : Lelej 1985: 194, ♂; 2002: 51, ♂; Lelej & Osten 2004: 255, ♂; Pagliano et al. 2020: 170.

Mutilla quinquedentata Morawitz, 1890: 638 , ♂, holotype, ♂, Repetek, 16.V.1889, on light, A. Semenov, probably lost. Junior subjective synonym of Mutilla dichroa Sichel et Radoszkowski, 1869 according to André, 1898: 6.

Ephutomma quinquedentata : Bischoff 1920: 149, 153, ♂ ( Iraq: Al Faw).

Ephutomma quinquedentata mesopotamica Bischoff, 1920: 149 , 153, ♂, holotype, ♂, Centr.- Mesopotamien, 518, Teklekberge [boundary between Syria and Turkey], 14.VII.[19]13, Exp. v. Oppenheim, Kohl S. // mesopotamica Bisch. det. Bischoff // Type // Dentilla dichroa (Sich.+Rad.) B. Petersen det. 1989 // Zool. Mus. Berlin [ MNHU], examined; Pagliano et al. 2020: 173 (as separate species). Syn. nov.

Smicromyrme sabulosa Skorikov, 1935: 313 , 317, ♀, holotype, ♀, Kerman, strana Megas, Shurab [Sistan & Baluchestan, Sor Ab, 27°10'24"N 60°41'17"E], 13–14.II.1901, N. Zarudny [ZIN], examined. Syn. nov.

Smicromyrme (Dentilla) sabulosa : Lelej & Kabakov 1980: 195, ♀.

Dentilla sabulosa : Lelej 1985: 191, 196, ♀; 2002: 52, ♀; Pagliano et al. 2020: 171.

Eremomyrme quinquedentata : Invrea 1965: 91, ♂ ( Israel).

Eremomyrme bicolor : Invrea 1965: 90, ♂ ( Israel, Palestine).

Diagnosis. MALE. Ocelli large, OOD 2 × diameter of lateral ocellus. Head reddish-brown. Wings hyaline with darkened apex. Anterior border of clypeus with narrow medial projection whose dorsal length is about equal to its width. Basal half of T2 with dense separate punctures. POD:OOD ratio 0.8 × and more, ocelli large. Upper carina of mandible high. Longitudinal carina between antennal tubercles and base of clypeus well developed. First subbasal tooth of the mandible with short carina much less than distance from the base of this carina to the base of the mandible. T7 apically with weak notch. T1 length less than its maximal width. Lower subbasal lobe of mandible invaginated below, height of subbasal tooth more than smallest distance between notch and upper carina of mandible. Body length 9.5–13.0 mm. FEMALE. T2 with basal medial spot of pale setae. T4–5 with pale setae. Eye located about same distance from articulation of mandible and posterior margin of vertex. Ratio of smallest distance between eyes to longitudinal eye diameter 1.6–1.8 ×. Length T2 0.8 × its maximal width. Body length 9.0–10.0 mm.

Material examined. (157♂ 3♀). Type material. Holotype of Ephutomma quinquedentata mesopotamica Bischoff, 1920 , ♂. Additional material. Turkmenistan: Dus-olum [south-western Turkmenistan], J. Pomerantsev / к. Моравица / 5-dentata F.Mor. 1♂ " [ ZIN]; GoogleMaps Peskimar, D. Glasunov [1893], 1♂ [ ZIN]; GoogleMaps Kopet-Dag, Firyuza, 8.VI.1903, K.Anger, 3♂ [ ZIN]; GoogleMaps 8, 20.VII.1991, on light, V. Dubatolov, 2♂ [ IBSS]; GoogleMaps East Kopet-Dag, Charlyk, 37°09ʹ45ʺN 59°49ʹ22ʺE, 26–27.VI.1991, V. Krivokhatsky, 1♂ [ IBSS]; GoogleMaps Bairam-Ali , 27.IX.1896, K.Anger, 1♂ [ ZIN]. GoogleMaps Afghanistan: (Uruzgan, Kabul), 2 ♂ ( Lelej & Kabakov 1980). Herat, Zulfagar [Sulfagar], [28.V.1893] D.Glasunov / к. Моравица / 5-dentata F.Mor. 3♂ " [ ZIN]. GoogleMaps Iran: (Hormozgan, Fars, Ilam), 100♂ ( Lelej & Osten 2004; Lelej et al. 2008) Tehran, 14.VIII.1951, N. Shutova, 1♂ [ ZIN]. GoogleMaps Razavi Khorasan Prov., Nerduali, [1893], D. Glasunov / к. Моравица / 5-dentata F.Mor. 2♂ " [ ZIN]; GoogleMaps South Khorasan Prov., Nehbandan, 24.VI–2.VII.1901, N. Zarudny, 1♂ [ ZIN]; GoogleMaps Yazd Prov., ca 23 km SW Kharanaq, 1600 m, on light, 32°11ʹN 54°30ʹE, A. Timokhov, 1♂ [ IBSS]. GoogleMaps Kerman Prov., 75 km S Sirjan, 9.IV.1970, Ressel, 1♀ [ ZSM]; GoogleMaps Keshi [Keshit], 28.III–2.IV.1901, N. Zarudny, 3♂ [ ZIN]; GoogleMaps 25 km NE Jiroft, 5 km E Saghdar, 2000–2400 m, on light, 28°50ʹN 57°56ʹE, A. Timokhov, 15♂ [ IBSS]. GoogleMaps Fars Prov., 50 km NNE of Shiraz, Bamoo Res., 29°45ʹN 52°46ʹE, 18–28.V.2000, Yu. Marusik, 2 ♂ 1♀ [ IBSS]; GoogleMaps Sistan & Baluchestan, Bampur, 14.V.1901, N. Zarudny, 1♂ [ ZIN]; GoogleMaps Karavandar [Karvandar], 23–24.IV. 1901, N. Zarudny, 4♂ [ ZIN]; GoogleMaps Sargad , 29–31.VIII.1898, N. Zarudny, 2♂ [ ZIN]; GoogleMaps Sargad , Chaashen-Sadk , 28–30.IV.1901, N. Zarudny, 1♂ [ ZIN]; GoogleMaps Shurab [Sor Ab], 27.VI.1898, N. Zarudny, 1♂ [ ZIN]; GoogleMaps Bazman , 28.VI–3.VII.1898, N. Zarudny, 3♂ [ ZIN]; GoogleMaps Makran , Rong River , 29.III.1901, N. Zarudny, 1♂ [ ZIN]; GoogleMaps Khash , 20.V.1955, D. Steinberg, 1♀ [ ZIN]. GoogleMaps Israel: 50 km SE Beer Sheva, Wadi En Aqrabbin , Negev , 30°57ʹN 35°08ʹE, 8.V.1996, Ch. Schmid-Egger, 1♂ [ IBSS]. GoogleMaps Jordan: Jemini , 19.XI.1966, coll. G. Pagliano, 1♂ [ IBSS]. GoogleMaps Oman: NE, S of Sur, 15 km W of Al Askharah, 13.III.2015, M. Snižek, 1♂ [ IBSS]. GoogleMaps Saudi Arabia: Riyadh, 1♂ [ OLML] GoogleMaps .

Distribution. Turkmenistan, Afghanistan, Iran, Iraq, Syria (new record), Jordan (new record), Israel, Palestine, Saudi Arabia (new record), Oman (new record), Egypt (Sinai).

Remarks. The holotype of Мutilla quinquedentata Morawitz, 1890 from Repetek is not found in the ZIN where the Morawitz collection is deposited. This holotype is probably lost. In the ZIN collection there are 7♂ from Nerduali, Sulfagar, Peskimar collected by D. Glasunov in the Transcaspian region and northern Iran and Dus-olum collected by J. Pomerantsev in south-western Turkmenistan, all are with Morawitz identification label " 5-dentata F.Mor., ♂ ". They cannot be syntypes ( Lelej 1985), because they were collected by D. Glasunov in 1893 (after the description of М. quinquedentata ) ( Semenov 1896). Børge Petersen (unpublished) labelled the holotype of Ephutomma quinquedentata mesopotamica Bischoff, 1920 , ♂ in MNHU as Dentilla dichroa ( Sichel et Radoszkowski, 1869) . As this synonymy has not yet been published, I hereby support it and propose it as a new synonymy.

The female of D. dichroa ( Sichel et Radoszkowski, 1869) may eventually be recognized as D. sabulosa ( Skorikov, 1935) due to its co-occurrence in Iran: Sistan and Baluchestan Prov. and especially Fars Prov., 50 km NNE of Shiraz, Bamoo Res., 29°45ʹN 52°46ʹE, 18–28.V.2000, Yu. Marusik, 1♀ and 2 ♂. There is no direct evidence (pair collected in copula) to support this relationship.