Acanthocnema pulchripennis, Sinclair, 2025

Acanthocnema pulchripennis sp. nov.

urn:lsid:zoobank.org:act:

Figs 29–34, 62

Diagnosis

This species is distinguished by the yellowish to yellowish-brown body, proepimeral seta present, fore femur with three long spine-like posteroventral setae, wing with discal crossband, without dark spot at apex of M

1

, and crossvein dm-m obliquely aligned with apex of M

4

.

Etymology

The specific epithet is derived from the Latin ‘ pulcher ’ (‘beauty’) and ‘ penna ’ (‘wing’), in reference to the unusually patterned wing.

Type material

Holotype

CANADA – Alberta • ♂; “CAN: AB: Peter Lougheed / PP; Boulton / Fox Ck Tr / 14.vii.2015; 1730 m / N 50°37.03′W 115°06.67′ / B.J. Sinclair; ex. creek/trail // CNC / 905982 View Materials // HOLOTYPE / Acanthocnema / pulchripennis / Sinclair [red label]”; CNC, CNC905982 View Materials .

Paratypes

CANADA – Alberta • 1 ♂; Moraine Lake ; [ 51°19′21″ N, 116°11′08″ W]; 3 Aug. 1923; J. McDunnough leg.; CNC GoogleMaps • 1 ♀; Waterton Lakes NP; [ 49°02′45″ N, 113°54′55″ W]; 1–5 Jul. 1980; H.J. Teskey leg.; CNC GoogleMaps . – British Columbia • 1 ♂, 1 ♀; Lisadele Lake ; 58°41′ N, 133°4′ W; 4800 ft a.s.l.; 10 Aug. 1960; W.W. Moss leg.; CNC GoogleMaps • 1 ♂; Manning PP, Cascade Lookout road; 20–21 Jul. 2015; 49°04.00′ N, 120°46.79′ W; 1240 m a.s.l.; B.J. Sinclair leg.; stream; CNC GoogleMaps • 1 ♂; Manning PP, Heather Trail to Buckhorn Camp ; 49°06.19′ N, 120°45.56′ W; 1785–1995 m a.s.l.; 21 Jul. 2015; B.J. Sinclair leg.; CNC GoogleMaps • 2 ♂♂, 1 ♀; Penticton, Apex Mt ; [ 49.390° N, 119.903° W]; 5800 ft a.s.l.; 13 Aug. 1967; J.R. Vockeroth leg.; CNC GoogleMaps • 1 ♂; same data as for preceding; KUMF GoogleMaps • 1 ♂; same data as for preceding; 5000 ft a.s.l.; 4 Aug. 1967; CNC GoogleMaps • 1 ♂; Silver Star PP, Sovereign Lake Rd ; 50°21.64′ N, 119°06.14′ W; 1665 m a.s.l.; 23 Jul. 2015; B.J. Sinclair leg.; CNC GoogleMaps • 1 ♂; Stagleap PP, Bridal Lake Trail ; 49°03.52′ N, 117°02.44′ W; 18 Jul. 2015; B.J. Sinclair leg.; CNC GoogleMaps • 1 ♂, 1 ♀; Vancouver Is, Helen Mackenzie Lake ; 48°42′ N, 125°19′ W; 1160 m a.s.l.; 27 Jul. 1994; H. Goulet leg.; wet meadow MT; CNC GoogleMaps . – Yukon • 1 ♀; Whitehorse, Wolf Ck, Cpgd ; [ 60.608° N, 134.942° W]; 17 Jun.–14 Jul. 1987; S.A. Marshall leg.; pits near spring; DEBU GoogleMaps .

Other material examined

CANADA – British Columbia • 1 ♂ (identification based on online image); Robson ; [ 49.337°N 117.665°W]; 24 Jul. 1958; H.R. Foxlee leg.; UBCZ GoogleMaps .

Description

Body yellowish to yellowish-brown ( Fig. 29), with dense white pruinescence. Body length: 5–6 mm; wing length: 4.5–5.3 mm.

Male

HEAD ( Fig. 29). Frontal vitta yellow to yellowish-brown anteriorly, matt; fronto-orbital plate brown, with numerous dark setulae. Chaetotaxy: anterior orbital seta proclinate, posterior 2 orbital setae lateroclinate; 3 inclinate frontals; 1 ocellar, 1 inner vertical, 1 outer vertical; postocellar setae weak, divergent; row of dark postocular setae; 1 vibrissa, 1–2 strong subvibrissae. Gena silvery pruinescent, depth ¼ eye height; postgenal setae white, soft, length subequal to width of postgena. Face, parafacial and occiput silvery pruinescent. Palpus yellowish white with long white setae. Proboscis with broad, shiny prementum, with scattered white setae, shorter than width of prementum. Antenna yellow, pedicel slightly darker; postpedicel rounded apically, 2 × as long as broad. Arista pubescent on basal third and bare apically.

THORAX ( Fig. 29). Yellowish-brown, darker along dorsocentral series; greyish vitta along acrostichal series. Acrostichals biserial, short, slender; 2+3 dorsocentrals; 1+2 intra-alars; 1+1 supra-alars; 1–2 postpronotals; 2 notopleurals, 2 postalars; scutellum with pair of strong apical setae and pair of strong basal setae; scutellar disc with a few scattered setulae. Proepisternal seta present. Proepimeral seta present. Anepisternum with dark setulae on posterior half and 1 long seta and several long thinner setae near posterior margin. Katepisternum with 1 strong, long seta in posterodorsal corner and numerous yellow setulae.

LEGS ( Fig. 30). Yellowish, gradually darker apically. Fore femur with 3 posteroventral spine-like setae on basal half, longer than width of femur; weakly defined row of anteroventral setae, stronger near mid-length; 4 posterodorsal setae on apical half, shorter than width of femur. Fore tibia with ventral apical spur, 1 anterodorsal slightly proximal to mid-length and 1 preapical dorsal seta; biserial row of short spinules ventrally. Fore tarsomere 1 with dense mat of anterior setulae. Mid femur with 4 long posteroventral setae on basal half, shorter than width of femur; row of 5–6 anterior setae; 1 preapical anterior seta and 1 posterior preapical seta. Mid tibia with 1 anterodorsal seta beyond mid-length and 1 preapical dorsal seta; 1 anteroventral seta and 1 posteroventral apical seta. Hind femur with row of 4–6 anterodorsal setae; several widely spaced posteroventral setae. Hind tibia with 3 anterodorsal setae; 1 posterodorsal seta near mid-length; 1 dorsal preapical seta; 1 anteroventral apical seta.

WING ( Fig. 31). Infuscate, with discal crossband from costa to M 4, without dark spot at apex of M 1, veins black; crossvein dm-m obliquely aligned with M 4; M 4 beyond cell dm strong to wing margin, one-third length of crossvein dm-m; CuA+CuP weakened but reaching wing margin. Calypteres and halter yellow.

ABDOMEN. Darker dorsally than thorax, with greyish pruinescence; with long erect setae on tergite 2 anteriorly. Sternites 3 and 4 expanded laterally with elongate setae ( Fig. 32). Sternite 5 broad, with pair of cylindrical lobes with long setae, apex slightly tapered and incurved ( Fig. 32). Terminalia ( Figs 33–34): Epandrium rounded, narrow with short setae. Cercus short, apical margin rounded in lateral view; cerci closely approximated medially, with setae longer than cercus. Surstylus digitiform, apex tapered, with wavy setae of various lengths, some longer than length of surstylus. Phallapodeme elongate, rod-shaped. Pregonite strongly recurved on distal half, with 1 short seta at mid-length; apex tapered to point with several short subapical setae. Postgonite gently arched with rounded apex. Ejaculatory apodeme flattened and expanded with tapered ejaculatory duct. Distiphallus narrow, tube-shaped with jagged inner margin.

Female

Similar to male except, abdomen darker, lacking long lateral setae on sternites 3 and 4; terminalia not dissected.

Distribution

This species is known from northwestern North America, recorded from Yukon to southern British Columbia ( Fig. 62).