Paraskusella sarawak, Tang & Cranston & Peter S., 2025

Paraskusella sarawak , new species

( Figs. 1A, C, E, F View Fig ; 2A–C, G–I View Fig ).

Material examined. Holotype male, slide mounted in Euparal, MALAYSIA: Sarawak, Gunung Mulu National Park, Lupar tributary of Melinau River , 75 m a.s.l., 04°1′48.8″N 114°49′4.4″E, 12 June 2023, light trap, coll. H.Q. Tang ( EJNU) GoogleMaps . Paratype male, as holotype ( ANIC), teneral male in alcohol, as holotype except 11 July 2024 ( SFC)

Etymology. From the type locality, Sarawak, the largest state of Malaysia on the island of Borneo. To be treated as a noun in apposition.

Description. Adult male (n = 2). Colour ( Fig. 1A View Fig ). Generally yellow green, thorax without distinct vittae, postnotum brown. Foreleg with darker proximal band on femur, and stronger darkening on distal femur and all of tibia and tarsus. Middle and hind leg with distal darker band on femora, tibiae yellow, all tarsomeres browner apically.

Total length 2.8–3.1 mm, wing length 1.35–1.45 mm.

Head. Frontal tubercles absent. Flagellomeres 1–12, 370–390; flagellomere 13, 575–590; AR 1.51–1.55. Palpomeres 1–5: 25–30; 20–25; 87–100; 90–105; 140–148, Pm3 and Pm4 slightly thickened. Temporals 6–8, uniserial. Clypeus with 9–10 setae.

Thorax with rounded scutum with weak median hump or smoothly rounded. Antepronotals 2–3; acrostichals 0; dorsocentrals comprising 4 anterior and 3–4 more posterior; prealars 2–3. Scutellum with 4 setae.

Wing ( Fig.1C View Fig ) appears ‘smoky’ with some darkening alongside veins, anal lobe weak. VR 1.17–1.25, Setation: R 0, R 1 0, R 4+5, 0, squama bare.

Legs ( Fig. 1A View Fig , 2A–C View Fig ). Fore tibia with round scale occupying half circumference. Spurs on mid and hind combs straight, 25–40 long. LR 1 2.04–2.08, LR 2 0.60–0.62, LR 3 0.77–0.82; BV 1 1.39–1.41, BV 2 4.19–4.20, BV 3 2.88–2.89; SV 1 1.37– 1.39, SV 2 3.65–3.78, SV 3 2.74–2.80.

Hypopygium ( Figs. 1E, F View Fig ; 2G–I View Fig ). Anal tergite bands separated medially to form incomplete V–shape, with two median anal tergite setae; 6–8 basal-lateral setae submarginal on tergite IX. Anal point ( Fig 1F View Fig , lateral view) spatulate, narrower in mid-section, 30–40 long from sclerotised base. Transverse sternapodeme broad, shallowly rounded anteriorly, with relatively strong oro-lateral projections. Phallapodeme indistinct. Gonocoxite c. 60 long, gonostylus c. 100 long. slender, inwardly curved, tapering to rounded point. Superior volsella apically crescent shaped, 35–40 long, without microtrichia, 2 medially-directed setae arising from distinct tubercle bases close to basal inner margin, with only 1 seta on outer margin. Median volsella ( Fig. 2H View Fig ) c. 40 long, with strong stem, medio-distally with simple setae, medially broadened and apically recurved and tapered distally, mostly not extending beyond distal apex of superior volsella. Inferior volsella globular apically and extending beyond gonocoxite apex, microtrichiose apicoventrally, apically with medially and dorsomedially-directed, simple setae, without differentiated posteriorly directed strong seta.

Female, pupa and larva unknown.

Remarks. Paraskusella sarawak , new species, conforms substantially to the morphological generic diagnosis for males of Paraskusella Cranston (2018) . The thorax with only slight medio-dorsal bulge at most, is consistent with previously described species excepting Paraskusella hawkei Cranston, 2018 in which a protuberance is distinct ( Cranston, 2018). The male of P. sarawak is separated from congeners by all the wing veins being bare.