Cybaeolus rastellus ( Roth, 1967 )

Cybaeolus rastellus ( Roth, 1967) View in CoL

( Figs 1A, B View Figure 1 , 2J, M View Figure 2 , 5 View Figure 5 , 6 View Figure 6 , 15 View Figure 15 , 18 View Figure 18 , 19D–F View Figure 19 , 20C, D View Figure 20 , 23 View Figure 23 , 24 View Figure 24 , 28B View Figure 28 )

Mevianes rastellus Roth 1967: 308 pl. 50, figs 7, 8 [male holotype, two males, and five female paratypes from Región de Los Lagos   GoogleMaps , Llanquihue Prov., Isla Calbuco, c. 42° 0 ʹ S, 73° 30 ʹ W (c. –41.79°, –73.15°) 21–28 Feb. 1963, H.J. McMillin and A. Archer, in AMNH, examined].

Cybaeolus rastellus, Lehtinen 1967: 226 View in CoL .

Cybaeolus cf. rastellus, Wheeler et al. 2017 View in CoL : fig. 5 (sequenced, voucher ARAMR-290, reexamined).

Diagnosis

Both sexes are similar to C. delfini by having a rastellum of thick setae on the cheliceral promargin (although sometimes with thinner setae, see Fig. 15B View Figure 15 ). Males of this species are also similar to those of C. delfini by having a fleshy median apophysis in the copulatory bulb ( Figs 15E, H View Figure 15 , 19D, E View Figure 19 ) and a two-pointed retrolateral patellar process. In C. rastellus the median apophysis is distally wider, and the patellar process is directed posteriorly ( Fig. 19E, F View Figure 19 ), instead of anteriorly as in C. delfini ( Fig. 19H, I View Figure 19 ). Females are similar to those of C. pusillus by having the copulatory openings anteriorly in the epigyne ( Figs 20C, D View Figure 20 , 23B–D View Figure 23 ) but can be distinguished by these being further apart from each other, the shorter and thicker copulatory ducts, describing a closer loop on the mesal margin of the spermathecae, six-shaped in ventral view, and the smaller spermathecae.

Description

Male (holotype, complemented with data from other specimens when not visible in the holotype). Total length 2.63. Carapace 1.06 long, 0.92 wide. Abdomen 1.64 long, 0.80 wide. Eye diameters and interdistances AME 0.05, ALE 0.08, PME 0.07, PLE 0.08; AME – AME 0.05, AME –ALE0.06, PME–PME 0.10,PME– PLE 0.09, ALE–PLE 0.02; MOQ length 0.16, anterior width 0.15, posterior width 0.24. Anterior eye row slightly procurved in anterior view, posterior row straight in dorsal view. Clypeus height 0.12 (2.4× diameter of AME). Carapace castaneous with indistinct darker border, darker thoracic V-shaped mark, cephalic region weakly setose. Chelicerae orange-brown, without visible stridulatory file (weak stridulatory file observed in some specimens, on external-posterior side). Endites yellow-brown, longer than wide, posterior lateral edge strongly protruding ( Fig. 15C View Figure 15 ). Labium light brown. Sternum castaneous, as wide as long. Opisthosoma distinctly elongated, dorsum: whitishgrey with three to four sepia chevrons posteriorly, mottled laterally, without setae expect on anterior dorsum; venter: mottled greyish-white with two indistinct longitudinal marks extending from epigastric furrow to base of spinnerets, darker around spinnerets. Palpal cymbium oval, copulatory bulb with small median apophysis ( Fig. 15E, H View Figure 15 ), tibial apophysis long and procurved with several long, strong, upwardly curved setae at base, patellar apophysis recumbent on retrolateral side, with two or three teeth ( Figs 15D View Figure 15 , 19E, F View Figure 19 ). Spinnerets in relatively compact group ( Fig. 6A View Figure 6 ), ALS pigmented, PLS and PMS unpigmented. Proximal segment of ALS 2.4 × length of distal segment; distal segment of PLS 0.24× length of proximal segment. PLS 1.1 × length of ALS. ALS total length 0.24; basal width 0.12, distal segment length 0.07, proximal segment length 0.17. PMS total length 0.13. PLS total length 0.26; distal segment length 0.05, proximal segment length 0.21. Spigot morphology: ALS with one major ampullate gland spigot flanked by nubbin, and 12 piriform gland spigots ( Fig. 6B View Figure 6 ). PMS with one minor ampullate gland spigot and three aciniform gland spigots ( Fig. 6C View Figure 6 ). PLS with five aciniform gland spigots ( Fig. 6D View Figure 6 ). Epiandrous spigots absent ( Fig. 6E View Figure 6 ). Legs orange-brown, all articles strongly annulated, lightly spined. Leg measurements (formula 1243): leg I (total 4.13) femur 1.19, patella 0.32, tibia 1.00, metatarsus 0.99, tarsus 0.63; leg II (4.07) 1.15, 0.35, 1.04, 0.92, 0.62; leg III (3.06) 1.04, 0.28, 0.68, 0.61, 0.45, leg IV (3.45) 1.05, 0.32, 0.81, 0.77, 0.48, palp (1.30) 0.44, 0.25, 0.22, -, 0.39. Leg formula 1243. Pattern of spines: Femora: I d0-1-0, p0-0-1; II d0-1-1 (or 0-1-0), p0-0-1 (or p0-1-1-0); III d0-0-1, p0-0-1. Patellae: I d1-0-1; II d1-0-1; III d1-0-1; IV d1-0- 1. Tibia: I d1-0-1, p0-0-1; II d1-0-1, p0-1-1; III d1-0-1, p0-0-1, v0-1-1; IV d1-0-1, v0-1-1. Metatarsus: III p0-1-1, v0-0-2, r0-0-1; IV p0-1-1, v0-2-2, r0-0-1.

Female (paratype). Total length 3.63. Carapace 1.13 long, 0.92 wide. Abdomen 2.28 long, 1.42 wide. Eye diameters and spacing: AME 0.05, ALE 0.07, PME 0.06, PLE 0.08; AME – AME 0.06, AME –ALE 0.06, PME–PME 0.09, PME–PLE 0.06, ALE–PLE 0.02; MOQ length 0.21, anterior width 0.16, posterior width 0.21. Clypeus height 0.13 (2.6× diameter of AME). Carapace cephalic region castaneous, shading to orange-brown laterally and posteriorly very faint radiating thoracic marks, weakly setose in cephalic region. Chelicerae length 0.47, ratio: 0.51, dark orange-brown, with prominent rastellum near base of fang. Endites short, as wide as long, without the posterior lateral extension of the male. Labium dusky. Sternum orange-brown, about as long as wide. Abdomen dorsum grey mottled with white, dark purplish-brown mark over cardiac region, dark band around posterior tip of abdomen, venter; without markings, sparsely setose along midline. Epigyne rugose, copulatory openings on shallow elevations ( Fig. 18G View Figure 18 ), copulatory duct begins with a loop anterior to the spermatheca, continues with a second loop, and connects through the ventral part of the spermatheca ( Figs 18H View Figure 18 , 20C, D View Figure 20 , 23B–D View Figure 23 ); accessory bulb as a small bump with gland ducts near the copulatory opening ( Fig. 18I View Figure 18 ). Spinnerets in relatively compact group ( Fig. 5A View Figure 5 ), ALS conical with wide base, distal segment of PLS very short, proximal segment of ALS 5.5 × length of distal segment; distal segment of PLS 0.19× length of proximal segment: PLS 1.2 × length of ALS. ALS total length 0.26; basal width 0.15, distal segment length 0.04, proximal segment length 0.22. PMS total length 0.20. PLS total length 0.31; distal segment length 0.05, proximal segment length 0.26. Spigot morphology: ALS with one major ampullate gland spigot flanked by a nubbin and 13 piriform gland spigots ( Fig. 5B View Figure 5 ). PMS with one minor ampullate gland spigot, two cylindrical gland spigots, and eight aciniform gland spigots ( Fig. 5C View Figure 5 ). PLS with one cylindrical gland spigot and 11 aciniform gland spigots ( Fig. 5D View Figure 5 ). Spinneret topology PLS 70 ° from ALS; PMS 45–50 View Materials ° anterior to ALS. Legs orange-brown, without distinct annulations but with dusky patches ventrally on femora and distally on metatarsus, pedipalp tarsi and tibiae dark orange-brown with patellae and femora a sharply contrasting light straw colour. Legs measurements: leg I (total 3.84), femur 1.11, patella 0.31, tibia 0.95, metatarsus 0.83, tarsus 0.64; leg II (3.86) 1.10, 0.37, 0.94, 0.90, 0.55; leg III (2.91) 0.90, 0.29, 0.64, 0.67, 0.41, leg IV (3.57) 1.06, 0.30, 0.86, 0.82, 0.53, palp (1.18), 0.33, 0.20, 0.27, -, 0.38. Pattern of spines: pedipalp patellae: d1-0- 1; tibiae: d1-0-0, p0-0-1; tarsi: d1-0-0, p1-1-1, v0-0-2, r1-1-1. Femora: I d0-0-1, p0-0-1 (or 2); II d0-0-1. Patellae all with: d1-0-1. Tibiae: I d1-0-1, p0-0-1; II d1-0-1, p0-0-1; III d1-0- 1, p0-0-1, vO1-1; IV d1-0-1, v0-1-1. Metatarsi: III and IV as male. Trichobothrial distribution tibiae: d1-2, metatarsi: d0-0- 1, tarsi: d0-1-0 or 0-1-1-0 ( Fig. 18D View Figure 18 ).

Variability and colour polymorphism

Some males have a weak stridulatory file on the external-posterior side of the chelicerae which is difficult to see.The cheliceral rastellum may have less conspicuous setae in some specimens. The body coloration is extremely variable ( Figs 2J, View Figure 2 M, 24 View Figure 24 ), even among specimens from the same locality and collection event ( Fig. 24A, D View Figure 24 ).

Natural history

Specimens were found in webs on foliage of trees, bamboo and shrubs in humid forests ( Fig. 1A, B View Figure 1 ).

Distribution

Known from forests in Chile from Región de Ñuble to Parque Nacional Río Simpson , in Región de Aysén, and in wet forests of adjacent Argentina in Parque Nacional Nahuel Huapi in Neuquén Province, and Parque Nacional Los Alerces in Chubut province ( Fig. 28B View Figure 28 ).

Other material examined

ARGENTINA: Chubut: Parque Nacional Los Alerces: Lago Futalaufquen [–42.845°, –71.705°], Jan. 1990, M. Ramírez, 1♂ ( MACN); Lago Menéndez, Pto Sagrario [–42.611°, –71.892°], 29 Nov. 1996, wet forest with alerce ( Fitzroya cupressoides ), M. Ramírez, 2♀ (MACN-Ar 42799). Neuquén: Parque Nacional Nahuel Huapi: Puerto Blest, Sendero de la Bahía, S 41.02583, W 71.82250, 789 m elev., 4 Dec. 2024, secondary forest, beating, A. Pérez-González, L.A. Martínez, M.J. Ramírez, and W.L. Porto, 1♀ (MACN-Ar 46685 MJR-2798); Lago Ortiz Basualdo [–40.983°, –71.864°], Jan. 1990, M. Ramírez, 1♂ 1♀ ( MACN); Puerto Blest [–41.033436, –71.816678], 10 Jan. 1998, M. Ramírez, 3♀ (MACN-Ar 21086); Puerto Blest [–41.033436, –71.816678], 7 Jan. 2000, L. Lopardo and A. Quiaglino, 1♂ (MACN-Ar 21087); PuertoBlest, SenderodelaBahía, 41.02583º S 71.82250º W, 789 m elev., 4 Dec. 2024, secondary forest, manual collecting, A. Pérez-González, L. Martínez, M. Ramírez, and W. Porto, 2♂, 2♀ (MACN-Ar 46861); Puerto Blest, trail to Laguna Los Cántaros, 41.013754º S, 71.822738º W, 820–890 m elev., 28 Dec. 2010, wet forest, M. Ramírez, V. Werenkraut, and S. Aisen, 1♂ (MACN-Ar 33597). Río Negro: Parque Nacional Nahuel Huapi: Lago Frías superior [–41.044°, –71.8°], 1 Feb. 1990, M. Ramírez, 1♂ 2♀ ( MACN); Puerto Blest, sector Lago Frías, trail before reaching Puerto Frías, 41.04056º S, 71.80083º W, 754 m elev., 5 Dec. 2023, secondary forest, beating, A. Pérez-González, L.A. Martínez, M.J. Ramírez, and W.L. Porto, 1♀ (MACN-Ar 46683 MJR-2794); trail to Co. La Mona, near Lago Espejo Chico, 40.580069º S, 71.702972º W, 890–940 m elev., 5 Jan. 2011, forest with Nothofagus dombeyi , Chusquea, M. Ramírez and V. Werenkraut , 1♀ 1 (MACN-Ar 34362 MLB-0942). CHILE: Región de Aysén: Parque Nacional Río Simpson: 22 km E Aisén [–45.562°, –72.325°], 300 m elev., 5 Feb. 1985, wet forest, N. Platnick and O. Francke, 1♀ ( AMNH); S margin of Río Simpson [–45.562°, –72.325°], 17 Feb. 1991, M. Ramírez, 2♂ 5♀ ( MACN). 15 km S Las Islas Juntas, 30 km Puyuhuapi [–44.09°, –72.446°], 100 m elev., 30 Dec. 1984 to 28 Jan. 1985, Nothofagus forest, S. and J. Peck, 5♂ 1♀ ( AMNH). 16 km W Cisnes Medio, Río Grande [–44.685°, –71.968°], 200 m elev., 30 Dec. 1984 to 28 Jan. 1985, mature beech forest, S. and J. Peck, 1♂ ♀ ( AMNH). Región de La Araucanía: Parque Nacional Nahuelbuta: 37º47 ʹ S, 73º00 ʹ W, 1200 m elev., 12 Feb. 2005, fogging Nothofagus antarctica, J.E. Barriga , 1♂ (MACN-Ar 43239); 1♂ (MACN-Ar 43240). Parque Nacional Ñielol: nr. Temuco [–38.723°, –72.587°], 250 m elev., 14–30 Dec. 1982, pyrethrin fogging tree bark, forest remnant/ Nothofagus, A. Newton and M. Thayer , 1♀ ( AMNH). 15 km NE Villarrica, Flor del Lago [–39.2°, –72.16°], 300 m elev., 14 Dec. 1984 to 10 Feb. 1985, Nothofagus forest, S. and J. Peck, 1♂ 2♀ ( AMNH). 17 km W Angol [–37.8°, –72.96°], 800 m elev., 8 Dec. 1984 to 16 Feb. 1985, Nothofagus forest, S. and J. Peck, 1♂ 1♀ ( AMNH). 21 km NE Pucón, Lago Caburga [–39.18°, –71.81°], 600 m elev., 15 Dec. 1984 to 10 Feb. 1985, mixed forest remnant, S. and J. Peck, 2♂ ♀ ( AMNH). Monumento Natural Contulmo: 38.01314º S, 73.18648º W, 341 m elev., 9 Jan. 2020, M. Ramírez, E. Soto, J. Wilson, and D. Poy, 1♀ (MACN-Ar 41385); 1♂ 2♀ (MACN-Ar 41388); 38º00 ʹ 46.8 ʹʹ S, 73º11 ʹ 15.4 ʹʹ W, 360 m elev., 10–11 Feb. 2005, M. Ramírez and F. Labarque, 1♀ (MACN-Ar 33625 MLB-4285); 3♀ (MACN-Ar 42801); 4♀ (MACN-Ar 42803); 3♀ (MACN-Ar 42805); Berlese funnel, 1♀ (MACN-Ar 43427); [–38.013°, –73.188°], 300 m elev., 11–12 Feb. 1992, N. Platnick, P. Goloboff, and M. Ramírez, 3♂ 5♀ ( AMNH); 2♂ 1♀ ( AMNH); 38º00 ʹ 46.2 ʹʹ S, 73º11 ʹ 15.4 ʹʹ W, 360 m elev., 12–13 Jul. 2010, manual night collecting, A. Ojangueren, L. Piacentini, E. Soto, and D. Valdivia, 1♀ (MACN-Ar 42822 AGB-0010); 1♂ (MACN-Ar 42865 LNP-4082); 1♀ (MACN-Ar 42873); 1♀ (MACN-Ar 42880 LNP-4077); general collecting, 2♀ (MACN-Ar 42872); [–38.014321, –73.180069], 17–21 Dec. 1998, M. Ramírez, L. Compagnucci, C. Grismado, and L. Lopardo, 1♀ (MACN-Ar 21090); 2♂ 2♀ (MACN-Ar 21088); [–38.027°, –73.203°], 12 Jan. 1989, M. Ramírez, 1♂ 5♀ ( MACN); sendero Lemu Mau, 38.01314º S, 73.18648º W, 341 m elev., 9 Feb. 2012, manual night collecting, M. Ramírez, M. Izquierdo, P. Michalik, C. Wirkner, and K. Huckstorf, 1♀ (MACN-Ar 35134). Región de Los Lagos: Chiloé , Isla Quinchao: 3.5 km from Curaco de Velez [–42.44°, –73.6°], 6 Feb. 1988, T. Cekalovic, 1♀ ( AMNH); Quetro [–42.418°, –73.58°], 16 Feb. 1996, T. Cekalovic, 1♀ ( AMNH). Isla Chiloé : Chepu [–42.04°, –73.97°], 17 m elev., 29 Nov. 1981, N. Platnick and R. Schuh, 1♂ ♀ ( AMNH); Chepu, NW coast Chiloé Island [–42.04°, –73.97°], 75 m elev., 21 Feb. 1992, N. Platnick, P. Goloboff, and M. Ramírez, 3♀ ( AMNH); km 1 N Puente Río Pudeto [–41.867°, –73.768°], 17 Feb. 1996, T. Cekalovic, 1♀ ( AMNH); km 1 W Huillinco [–42.68°, –73.91°], 24 May 1998, T. Cekalovic, 1♀ ( AMNH); Lago Huillinco, N margin, 4.6 km (air) ESE Cucao, 42.64117º S, 74.04763º W, 12 m elev., 16 Feb. 2012, wet forest, M. Ramírez, M. Izquierdo, P. Michalik, C. Wirkner, and K. Huckstorf, 3♂ 1♀ (MACN-Ar 34138); Pío-Pío [–42.814°, –73.584°], 10–12 Mar. 1987, L. Peña, 2♀ ( AMNH); Piruquina [–42.38°, –73.79°], 19 Feb. 1983, T. Cekalovic, 1♀ ( AMNH); Río Cipresal, 42º 35.181 ʹ S, 74º 05.576 ʹ W, 2 Mar. 2008, Fogging 150cc/l N. nitida , 1♀ ( CASENT 9034054 JMA-0223). Parque Nacional Chiloé : 15 km al S Chepu [–42.14°, –74.05°], 3 Feb. 1991, M. Ramírez, 3♂ 7♀ (MACN-Ar 21085); Arroyo Cole Cole 25 km N Cucao [–42.406°, –74.108°], 8–11 Feb. 1991, M. Ramírez, 4♂ 8♀ ( MACN). Llanquihue, Caleta La Arena [–41.681°, –72.654°], 30 Jan. 1990, M. Ramírez, 1♀ ( MACN). Llanquihue, Ensenada [–41.179°, –72.553°], 50 m elev., 16 Mar. 1965, H.W. Levi, 1♂ ( MCZ). Parque Nacional Alerce Andino: 22–24 Jan. 1997, L. Compagnucci, 1♂ 3♀ (together with one immature) (MACN-Ar 21084); sector Sargazo, between start sendero Los Ulmos and sendero Laguna Sargazo, 41.509747° S, 72.616501° W, 347 m elev., 3–4 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ ♀ (MACN-Ar 43953); sector Sargazo, sendero Laguna Sargazo, 41.51613° S, 72.604081° W, 382 m elev., 3 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♀ (MACN-Ar 43958); sector Sargazo, sendero Los Ulmos, 41.507524° S, 72.619967° W, 382 m elev., 3 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 3♂ 2♀ (MACN-Ar 43968); sendero Rodal del Alerce, 41.510801° S, 72.609275° W, 388 m elev., 4 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ ♀ (MACN-Ar 43959); sendero ‘Rodal Alerce’, 41º30 ʹ 35.8 ʹʹ S, 72º36 ʹ 44.6 ʹʹ W, 340 m elev., 4 Feb. 2005, wet forest, M. Ramírez and F. Labarque, 1♀ (MACN-Ar 33406); 1♀ (MACN-Ar 33407 MLB-4273); 5♂ 18♀ (MACN-Ar 42800); sector Correntoso, sendero ‘Huillifoten’, wet forest, 41º27 ʹ 53 ʹʹ S, 72º38 ʹ 43.4 ʹʹ W, 135 m elev., 3 Feb. 2005, M. Ramírez and F. Labarque, 1♀ (MACN-Ar 33382 MLB-4252); 1♀ (MACN-Ar 42797 ARAMR 000290); 1♂ 3♀ (MACN-Ar 42809); 1♀ (MACN-Ar 42807); trail to Saltos del Río Chaica 7–10 km NE Lenca, 41º34 ʹ 41.9 ʹʹ S, 72º33 ʹ 24.2 ʹʹ W, 205 m elev., 2 Feb. 2005, M. Ramírez and F. Labarque, 1♂ 1♀ (MACN-Ar 33393 male MLB-4261, female MLB-4262); 1♀ (MACN-Ar 10836); 1♀ (MACN-Ar 42798 ARAMR 000291); 6♂ 14♀ (MACN-Ar 42802 AGB-0019); 1♀ (MACN-Ar 42804 ARAMR 000292); 3♀ (MACN-Ar 42806). Parque Nacional Puyehue: 600 m elev., 12–20 Feb. 1979, L. Peña, 1♀ ( AMNH); 4.1 km E Anticura [–40.66°, –72.19°], 430 m elev., 19–26 Dec. 1982, trap site 662, Valdivian rain forest, A. Newton and M. Thayer, 18♀ ( AMNH); 26 Dec. 1982, Valdivian rain forest, A. Newton and M. Thayer, 1♀ ( AMNH); 8–26 Dec. 1982, 6♂ ♀ ( AMNH); Aguas Calientes, 40°44 ʹ S, 72°18 ʹ W, 500 m elev., 16 Jan. 1995, N. Platnick, K. Catley and D. Silva, 3♂ 6♀ ( AMNH); [–40.733°, –72.3°], 600 m elev., 18 Dec. 1984 to 8 Feb. 1985, malaise, Nothofagus forest, S. and J. Peck, 2♂ ♀ ( AMNH); 40°44 ʹ S, 75°18 ʹ 45 ʹʹ W, 450 m elev., J. Miller, I. Agnarsson, F. Alvarez, J. Coddington, and G. Hormiga, 12 Nov. 2000 to 2 Jan. 2001, 1♂ 1♀ ( USNM); 500 m elev., 2–5 Jan. 1982, L. Peña, 1♂ 1♀ ( AMNH); 450 m elev., 27 Jan. 1986, wet forest, N. Platnick and R. Schuh, 1♀ (together with one immature) ( AMNH); 430 m elev., 28 Jan. 28, 1986, primary forest, N. Platnick and R. Schuh, 1♀ ( AMNH); 1♂ 1♀ ( AMNH); 425 m elev., 31 Jan. 1985, Valdivian forest, N. Platnick, and O. Francke, 2♀ ( AMNH); 13–17 Dec. 1998, M. Ramírez, L. Compagnucci, C. Grismado, and L. Lopardo, 1♂ (MACN-Ar 21091); 7♂ 12♀ (MACN-Ar 42796); Aguas Calientes, Derumbes forest trail [–40.733°, –72.3°], 500 m elev., 20 Dec. 1984 to 8 Feb. 1985, S. and J. Peck, 5♂ ♀ ( AMNH); Anticura, 40º39 ʹ 58.6 ʹʹ S, 72º10 ʹ 24 ʹʹ W, 355 m elev., 5 Feb. 2005, M. Ramírez and F. Labarque, 5♀ (MACN-Ar 42808); Antillanca road [–40.776°, –72.209°], 470–720 m elev., 18–24 Dec. 1982, pyrethrin fogging mossy log: 655 m, Valdivian rain forest, A. Newton and M. Thayer, 1♀ ( AMNH); sector Aguas Calientes, sendero Berlin, 40.741952° S, 72.299176° W, 490 m elev., 10 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ 2♀ (MACN-Ar 43966); sector Aguas Calientes, sendero El Recodo, 40º44 ʹ 17.1 ʹʹ S, 72º18 ʹ 25.6 ʹʹ W, 367 m elev., 10 Jul. 2010, Beating, A. Ojangueren, L.Piacentini and E.Soto, 1♀ (MACN-Ar 42820 LNP-4131); beating, 1♂ (MACN-Ar 42821 LNP-4132); 1♀ (MACN-Ar 42823); 2♀ (MACN-Ar 42870); sector Aguas Calientes, sendero Los Pioneros, 40.73794° S, 72.311679° W, 572 m elev., 9 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ 6♀ (MACN-Ar 43969); sector Aguas Calientes, sendero Recoba, 40.735181° S, 72.30985° W, 474 m elev., 10 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ 2♀ (MACN-Ar 43954); sector Anticura, sendero Chile, 40.670723° S, 72.163633° W, 457 m elev., 11 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ ♀ (MACN-Ar 43960); sector Anticura, sendero Pudú, 39º44 ʹ 08.2 ʹʹ S, 71º40 ʹ 13 ʹʹ E, 396 m elev., 8 Jul. 2010, under logs, A. Ojanguren, L. Piacentini, and E. Soto, 1♀ (MACN-Ar 42867 LNP-4141); manual night collecting, 1♀ (MACN-Ar 42878); sector Anticura, sendero Salto del Indio, 40.663337° S, 72.17616° W, 358 m elev., 11 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 3♂ ♀ (MACN-Ar 43964); sector Anticura, sendero Salto del Pudú, 40.675656° S, 72.164139° W, 480 m elev., 11 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ 1♀ (MACN-Ar 43973). 1 km E Termas de Puyehue [–40.66°, –72.2°], 305 m elev., 31 Jan. 1985, wet forest, N. Platnick and O. Francke, 1♀ ( AMNH). 25–27 km N Chaitén [–42.72°, –72.8°], 40 m elev., 17 Jan.1986, wet virgin forest, N. Platnick, P. Goloboff and R. Schuh, 1♀ ( AMNH). 36 km W La Unión [–40.18°, –73.46°], 600 m elev., 25–28 Mar. 1981, L. Peña, 2♀ ( AMNH). 4.5 km W La Unión, E El Mirador [–40.3°, –73.14°], 300 m elev., 1–2 Mar. 1987, L. Peña, 1♂ 1♀ ( AMNH). 70 km S Chaitén [–43.42°, –72.36°], 500 m elev., 16 Jan. 1986, wet streambank, N. Platnick, P. Goloboff, and R. Schuh, 1♀ ( AMNH). Anticura, E of Puyehue [–40.658°, –72.253°], 26–31 Aug. and 1–5 Sep. 1983, L. Peña, 1♀ ( AMNH). Hills S Maicolpue [–40.62°, –73.74°], 50 m elev., 19 Feb. 1992, N. Platnick, P. Goloboff, and M. Ramírez, 3♀ ( AMNH). Lago Chapo, 34 km E Puerto Montt [–41.42°, –72.56°], 300 m elev., 24 Dec. 1984 to 2 Feb. 1985, 2nd grow Nothofagus , S. and J. Peck, 4♂ 2♀ ( AMNH). Pucatrihue coast [–40.54°, –73.71°], 1–10 Feb. 1990, L. Peña, 1♀ ( AMNH). Termas de Puyehue [–40.736°, –72.308°], 240 m elev., 14 Mar. 1965, beach vegetation, H.W. Levi, 1♀ ( MCZ); 180 m elev., 24 Nov. 1981, forest, N. Platnick and R. Schuh, 1♀ ( AMNH); 460 m elev., 25 Nov. 1981, litter from moss and forest floor, N. Platnick and R. Schuh, 2♀ ( AMNH). Región de Los Ríos: Parque Nacional Alerce Costero: sendero Los Melies, 39.951827° S, 73.55019° W, 32 m elev., 5 Jan. 2023, M.P. Pacheco, L. Piacentini, and E.M. Soto, 1♂ ♀ (MACN-Ar 43962). 34 km WNW La Unión [–40.21°, –73.45°], 17 Dec. 1984 to 7 Feb. 1985, mixed evergreen forest, S. and J. Peck, 1♀ ( AMNH). 1 before instersection of Cariñanco to Parque Oncol [–39.7°, –73.355°], 12 May 2001, T. Cekalovic, 1♂ ♀ ( AMNH). Las Lajas, W La Unión, 9–13 May 1990, L. Peña, 4♀ ( AMNH); Las Trancas, W La Unión, Jun. 1990, L. Peña, 1♂ ♀ ( AMNH); 500 m elev., 6–10 Feb. 1988, L. Peña, 1♂ 3♀ ( AMNH). Parque Oncol: Quitaqui trail [–39.7°, –73.32°], 19 May 2001, T. Cekalovic, 2♂ 1♀ ( AMNH). Región de Ñuble: 9 km W San Fabian de Alico [–36.53°, –71.61°], 24 Feb. 1992, N. Platnick, P. Goloboff, and M. Ramírez, 1♀ ( AMNH). Región del Biobío: Arauco, Hualpén [–36.798°, –73.163°], 11 Jan. 1989, M. Ramírez, 1♀ ( MACN). Bosque de Ramuntcho [–36.752°, –73.185°], 14–16 Nov. 1961, A.F. Archer, 1♀ ( AMNH). Caramávida [–37.692°, –73.353°], 1–10 Jan. 1954, L.E. Pena, 3♀ ( CAS). Curiman [–36.95°, –73.13°], 29 Dec. 1994, T. Cekalovic, 1♀ ( AMNH). Estero Nonguén [–36.87°, –72.98°], 10 Oct. 1996, T. Cekalovic, 1♀ ( AMNH); 11 Dec. 1977, 1♀ ( AMNH). Lomas Coloradas [–36.9°, –73.12°], 27 May 1988, T. Cekalovic, 1♂ ♀ ( AMNH). Pata de Gallina [–38.07°, –73.23°], 560 m elev., 11 Feb. 1992, N. Platnick, P. Goloboff, and M. Ramírez, 1♂ 1♀ ( AMNH); 2♀ ( AMNH). Quebrada Caramávida, San Alfonso, reserva Arauco, 20 km ESE Antiguala, 37.70942º S, 73.17107º W, 750 m elev., 15 Jan. 2018, forest with Araucaria , general collecting, M. Ramírez, A. Ojanguren, A. Pérez Gonzalez, G. Azevedo, and W. Porto, 1♂ (MACN-Ar 43342 MJR-2264).