Onthophagus cyanellus, Bates, 1887

KEY TO THE ONTHOPHAGUS CYANELLUS View in CoL SPECIES COMPLEX AND CLOSELY RELATED SPECIES (modified from Zunino and Halffter 1988)

1. Pronotal prominence of major male trapezoidal ................................................ 2

1´. Pronotal prominence of major male triangular ..................................................... 3

2. Frontal carina of male raised laterally into 2 triangular tubercles ( Fig. 3 View Figs ). Apex of parameres not notched ( Fig. 8 View Figs ). Mexico to Panama ( Fig. 10 View Fig ) ............ O. cyanellus Bates, 1887

2´. Frontal carina of male raised laterally into 2 strongly developed horns ( Fig. 2 View Figs ). Apex of parameres notched ( Fig. 6 View Figs ). Central Highlands of Chiapas, Mexico ( Fig. 10 View Fig ) .................... ....... O. sancristobalensis Moctezuma and Halffter , new species

3. Pronotal punctures umbilicate. Central Highlands of Chiapas, Mexico ( Fig. 10 View Fig ) ................ ... O. chiapanecus Zunino and Halffter, 1988

3´. Pronotal punctures not umbilicate ........... 4

4. Dark blue dorsally. Head and pronotum shiny, contrasting strongly with dull elytra. Sierra Madre del Sur, Mexico ( Fig. 10 View Fig ) ............... ................... O. semiopacus Harold, 1869

4´. Dark blue-green dorsally, with microsculpture on head, pronotum, and elytra. El Salvador ( Fig. 10 View Fig ) ........................ O. salvadorensis Zunino and Halffter, 1988